[1] 王义平, 吴鸿, 徐华潮. 以昆虫作为指示生物评估森林健康的生物与生态学基础[J]. 应用生态学报, 2008, 19(7):1625-1630.

WANG Yiping, WU Hong, XU Huachao. Biological and ecological bases of using insect as a bio-indicator to asses forest health[J]. Chin J Appl Ecol, 2008, 19(7):1625-1630.
[2] 邢韶华, 姬文元, 郭宁, 等. 森林生态系统健康研究进展[J]. 生态学杂志, 2009, 28(10):2102-2106.

XING Shaohua, JI Wenyuan, GUO Ning, et al. Forest ecosystem health:its research progress[J]. Chin J Ecol, 2009, 28(10):2102-2106.
[3] 王义平, 吴鸿, 徐华潮. 浙江重点生态地区蝶类生物多样性及其森林生态系统健康评价[J]. 生态学报, 2008, 28(11):5259-5269.

WANG Yiping, WU Hong, XU Huachao. Butterfly diversity and evaluation of forest eco-system health in important ecological areas of Zhejiang Province[J]. Acta Ecol Sin, 2008, 28(11):5259-5269.
[4] 高光彩, 付必谦. 步甲作为指示生物的研究进展[J]. 昆虫知识, 2009, 46(2):216-222.

GAO Guangcai, FU Biqian. Advances of researches on carabid beetles as bioindicators[J]. Chin Bull Entomol, 2009, 46(2):216-222.
[5] 郭瑞, 王义平, 吴鸿. 森林凋落物层的节肢动物与森林健康的关系[J]. 林业科学, 2012, 48(3):122-127.

GUO Rui, WANG Yiping, WU Hong. Relationship between arthropods in forest litter and forest health[J]. Sci Silv Sin, 2012, 48(3):122-127.
[6] WEAVER J C. Indicator species and scale of observation[J]. Conserv Biol, 1995, 9(4):939-942.
[7] JUDAS M, DORNIEDEN K, STROTHMANN U. Distribution patterns of carabid beetle species at the landscape level[J]. J Biogeogr, 2002, 29(4):491-508.
[8] ANTVOGEL H, BONN A. Environmental parameters and microspatial distribution of insects:a case study of carabids in an alluvial forest[J]. Ecography, 2001, 24(4):470-482.
[9] MOLNAR T, MAGURA T, TÓTHMÉRÉSZ B, et al. Ground beetles (Carabidae) and edge effect in oak-hornbeam forest and grassland transects[J]. Eur J Soil Biol, 2001, 37(4):297-300.
[10] TYLER G. The ground beetle fauna (Coleoptera:Carabidae) of abandoned fields, as related to plant cover, previous management and succession stage[J]. Biodiversity Conserv, 2008, 17(1):155-172.
[11] KOIVULA M, PUNTTILA P, HAILA Y, et al. Leaf litter and the small-scale distribution of carabid beetles (Coleoptera, Carabidae) in the boreal forest[J]. Ecography, 1999, 22(4):424-435.
[12] 王建芳, 王新谱, 李秀敏, 等. 不同土地利用方式对河北坝上步甲物种多样性的影响[J]. 昆虫学报, 2010, 53(10):1127-1134.

WANG Jianfang, WANG Xinpu, LI Xiumin, et al. Influence of land uses on species diversity of carabid beetles (Coleopeta, Carabidae) in Bashang region, Hebei, Northern China[J]. Acta Entomol Sin, 2010, 53(10):1127-1134.
[13] 顾伟, 马玲, 刘哲强, 等. 凉水自然保护区不同林型地表甲虫物种多样性[J]. 东北林业大学学报, 2014, 42(1):131-135.

GU Wei, MA Ling, LIU Zheqiang, et al. Diversity of ground-dwelling beetle with different forest type in Liangshui Nature Reserve[J]. J Northeast For Univ, 2014, 42(1):131-135.
[14] 李明华. 清凉峰自然保护区志[M]. 长春:吉林人民出版社. 2006.
[15] 张希彪, 上官周平. 人为干扰对黄土高原子午岭油松人工林土壤物理性质的影响[J]. 生态学报, 2006, 26(11):3685-3695.

ZHANG Xibiao, SHANGGUAN Zhouping. Effect of human-induced disturbance on physical properties of soil in artificial Pinus tabulaeformis Carr. forests of the Loess Plateau[J]. Acta Ecol Sin, 2006, 26(11):3685-3695.
[16] BUTTERFIELD J. Carabid community succession during the forestry cycle in conifer plantations[J]. Ecography, 1997, 20(6):614-625.
[17] RAINIO J, NIEMEL J. Ground beetles (Coleoptera:Carabidae) as bioindicators[J]. Biodiversity Conserv, 2003, 12(3):487-506.
[18] SROKA K, FINCH O D. Ground beetle diversity in ancient woodland remnants in north-western Germany (Coleoptera, Carabidae)[J]. J Insect Conserv, 2006, 10(4):335-350.
[19] 于晓东, 周红章, 罗天宏. 神农架保护区大步甲和蜗步甲属生境选择与物种多样性研究[J]. 生物多样性, 2001, 9(3):214-221.

YU Xiaodong, ZHOU Hongzhang, LUO Tianhong. Habitat distribution and species diversity of Carabus and Cychrus in Shennongjia Nature Reserve, Hubei Province[J]. Biodiversity Sci, 2001, 9(3):214-221.
[20] LKEDA H, TSUCHIYA Y, NAGATA N, et al. Altitudinal life-cycle and body-size variation in ground beetles of the genus Carabus (subgenus Ohomopterus) in relation to the temperature conditions and prey earthworms[J]. Pedobiologia, 2012, 55(2):67-73.