[1] ROGERS J M, BULYK M L. Diversification of transcription factor-DNA interactions and the evolution of gene regulatory networks[J]. Wiley Interdisciplinary Reviews. Systems Biology and Medicine, 2018, 10(5): e1423. DOI: 10.1002/wsbm.1423.
[2] AHMAD MERAJ T, FU Jingye, ALI RAZA M, et al. Transcriptional factors regulate plant stress responses through mediating secondary metabolism[J]. Genes, 2020, 11(4): 346. DOI: 10.3390/genes11040346.
[3] WANG Yutao, CAO Yu, QIN Genji. Multifaceted roles of TCP transcription factors in fate determination[J]. New Phytologist, 2025, 245(1): 95−101. DOI: 10.1111/nph.20188.
[4] LIU Donghai, LUO Yin, HAN Han, et al. Genome-wide analysis of Citrus TCP transcription factors and their responses to abiotic stresses[J]. BMC Plant Biology, 2022, 22(1): 325. DOI: 10.1186/s12870-022-03709-3.
[5] KOSUGI S, OHASHI Y. PCF1 and PCF2 specifically bind to cis elements in the rice proliferating cell nuclear antigen gene[J]. The Plant Cell, 1997, 9(9): 1607−1619. DOI: 10.1105/tpc.9.9.1607.
[6] ZHOU Haiying, HWARARI D, MA Hongyu, et al. Genomic survey of TCP transcription factors in plants: phylogenomics, evolution and their biology[J]. Frontiers in Genetics, 2022, 13: 1060546. DOI: 10.3389/fgene.2022.1060546.
[7] VIOLA I L, ALEM A L, JURE R M, et al. Physiological roles and mechanisms of action of class Ⅰ TCP transcription factors[J]. International Journal of Molecular Sciences, 2023, 24(6): 5437. DOI: 10.3390/ijms24065437.
[8] CRAWFORD B C W, NATH U, CARPENTER R, et al. CINCINNATA controls both cell differentiation and growth in petal lobes and leaves of Antirrhinum[J]. Plant Physiology, 2004, 135(1): 244−253. DOI: 10.1104/pp.103.036368.
[9] DANISMAN S, VAN DER WAL F, DHONDT S, et al. Arabidopsis Class Ⅰ and class Ⅱ TCP transcription factors regulate jasmonic acid metabolism and leaf development antagonistically[J]. Plant Physiology, 2012, 159(4): 1511−1523. DOI: 10.1104/pp.112.200303.
[10] CHAHEL A A, ZENG Shaohua, YOUSAF Z, et al. Plant-specific transcription factor LrTCP4 enhances secondary metabolite biosynthesis in Lycium ruthenicum hairy roots[J]. Plant Cell, Tissue and Organ Culture (PCTOC), 2019, 136(2): 323−337. DOI: 10.1007/s11240-018-1518-2.
[11] RIECHMANN J L, HEARD J, MARTIN G, et al. Arabidopsis transcription factors: genome-wide comparative analysis among eukaryotes[J]. Science, 2000, 290(5499): 2105−2110. DOI: 10.1126/science.290.5499.2105.
[12] YAO Xuan, MA Hong, WANG Jian, et al. Genome-wide comparative analysis and expression pattern of TCP gene families in Arabidopsis thaliana and Oryza sativa[J]. Journal of Integrative Plant Biology, 2007, 49(6): 885−897. DOI: 10.1111/j.1744-7909.2007.00509.x.
[13] PARAPUNOVA V, BUSSCHER M, BUSSCHER-LANGE J, et al. Identification, cloning and characterization of the tomato TCP transcription factor family[J]. BMC Plant Biology, 2014, 14(1): 157. DOI: 10.1186/1471-2229-14-157.
[14] MA Xiaodong, MA Jianchao, FAN Di, et al. Genome-wide identification of TCP family transcription factors from Populus euphratica and their involvement in leaf shape regulation[J]. Scientific Reports, 2016, 6: 32795. DOI: 10.1038/srep32795.
[15] SHANG Xiaowen, HAN Zhaolan, ZHANG Dayan, et al. Genome-wide analysis of the TCP gene family and their expression pattern analysis in tea plant (Camellia sinensis)[J]. Frontiers in Plant Science, 2022, 13: 840350. DOI: 10.3389/fpls.2022.840350.
[16] ZHANG Jianping, TIAN Xinhui, YANG Yongxun, et al. Gleditsia species: an ethnomedical, phytochemical and pharmacological review[J]. Journal of Ethnopharmacology, 2016, 178: 155−171. DOI: 10.1016/j.jep.2015.11.044.
[17] LEE J, YI Jinmu, KIM H, et al. Cytochalasin H, an active anti-angiogenic constituent of the ethanol extract of Gleditsia sinensis thorns[J]. Biological and Pharmaceutical Bulletin, 2014, 37(1): 6−12. DOI: 10.1248/bpb.b13-00318.
[18] XIAO Dandan, LIU Jiahao, WANG Jing, et al. Chromosome-level de novo genome unveils the evolution of Gleditsia sinensis and thorns development[J]. Genomics, 2025, 117(2): 111004. DOI: 10.1016/j.ygeno.2025.111004.
[19] XIAO Feng, ZHAO Yang, WANG Xiurong, et al. Comparative transcriptome analysis of Gleditsia sinensis thorns at different stages of development[J]. Plants, 2023, 12(7): 1456. DOI: 10.3390/plants12071456.
[20] CAMOIRANO A, ARCE A L, ARIEL F D, et al. Class I TCP transcription factors regulate trichome branching and cuticle development in Arabidopsis[J]. Journal of Experimental Botany, 2020, 71(18): 5438−5453. DOI: 10.1093/jxb/eraa257.
[21] GASTALDI V, NICOLAS M, MUÑOZ-GASCA A, et al. Class Ⅰ TCP transcription factors TCP14 and TCP15 promote axillary branching in Arabidopsis by counteracting the action of Class Ⅱ TCP BRANCHED1[J]. New Phytologist, 2024, 243(5): 1810−1822. DOI: 10.1111/nph.19950.
[22] WANG Shenhao, YANG Xueyong, XU Mengnan, et al. A rare SNP identified a TCP transcription factor essential for tendril development in cucumber[J]. Molecular Plant, 2015, 8(12): 1795−1808. DOI: 10.1016/j.molp.2015.10.005.
[23] ZHANG Shengkui, LI Can, CUI Baihui, et al. TCP23-WRKY15 module negatively regulates lignin deposition and xylem development of wood formation in Populus[J]. International Journal of Biological Macromolecules, 2025, 306: 141656. DOI: 10.1016/j.ijbiomac.2025.141656.
[24] CHEN Chengjie, CHEN Hao, ZHANG Yi, et al. TBtools: an integrative toolkit developed for interactive analyses of big biological data[J]. Molecular Plant, 2020, 13(8): 1194−1202. DOI: 10.1016/j.molp.2020.06.009.
[25] LIVAK K J, SCHMITTGEN T D. Analysis of relative gene expression data using real-time quantitative PCR and the 2−ΔΔCT method[J]. Methods, 2001, 25(4): 402−408. DOI: 10.1006/meth.2001.1262.
[26] JIAN Bo, LIU Bin, BI Yurong, et al. Validation of internal control for gene expression study in soybean by quantitative real-time PCR[J]. BMC Molecular Biology, 2008, 9(1): 59. DOI: 10.1186/1471-2199-9-59.
[27] FENG Zhijuan, XU Shengchun, LIU Na, et al. Soybean TCP transcription factors: evolution, classification, protein interaction and stress and hormone responsiveness[J]. Plant Physiology and Biochemistry, 2018, 127: 129−142. DOI: 10.1016/j.plaphy.2018.03.020.
[28] CHALLA K R, AGGARWAL P, NATH U. Activation of YUCCA5 by the transcription factor TCP4 integrates developmental and environmental signals to promote hypocotyl elongation in Arabidopsis[J]. The Plant Cell, 2016, 28(9): 2117−2130. DOI: 10.1105/tpc.16.00360.
[29] WANG Zhishuo, WANG Ying, KOHALMI S E, et al. Squamosa promoter binding protein-like 2 controls floral organ development and plant fertility by activating asymmetric leaves 2 in Arabidopsis thaliana[J]. Plant Molecular Biology, 2016, 92(6): 661−674. DOI: 10.1007/s11103-016-0536-x.
[30] MARTÍN-TRILLO M, CUBAS P. TCP genes: a family snapshot ten years later[J]. Trends in Plant Science, 2010, 15(1): 31−39. DOI: 10.1016/j.tplants.2009.11.003.
[31] AGUILAR-MARTÍNEZ J A, POZA-CARRIÓN C, CUBAS P. Arabidopsis BRANCHED1 acts as in integrator of branching signals within axillary buds[J]. The Plant Cell, 2007, 19(2): 458−472. DOI: 10.1105/tpc.106.048934.
[32] WANG Han, MAO Yanfei, YANG Jun, et al. TCP24 modulates secondary cell wall thickening and anther Endothecium development[J]. Frontiers in Plant Science, 2015, 6: 436. DOI: 10.3389/fpls.2015.00436.
[33] KIEFFER M, MASTER V, WAITES R, et al. TCP14 and TCP15 affect internode length and leaf shape in Arabidopsis[J]. The Plant Journal, 2011, 68(1): 147−158. DOI: 10.1111/j.1365-313x.2011.04674.x.