[1] 白永飞, 黄建辉, 郑淑霞, 等. 草地和荒漠生态系统服务功能的形成与调控机制[J]. 植物生态学报, 2014, 38(2): 93−102.

BAI Yongfei, HUANG Jianhui, ZHENG Shuxia, et al. Drivers and regulating mechanisms of grassland and desert ecosystem services[J]. Chinese Journal of Plant Ecology, 2014, 38(2): 93−102. DOI: 10.3724/SP.J.1258.2014.00009.
[2] BAI Yongfei, COTRUFO M F. Grassland soil carbon sequestration: current understanding, challenges, and solutions[J]. Science, 2022, 377(6606): 603−608. DOI: 10.1126/science.abo2380.
[3] 张扬建, 朱军涛, 沈若楠, 等. 放牧对草地生态系统影响的研究进展[J]. 植物生态学报, 2020, 44(5): 553−564.

ZHANG Yangjian, ZHU Juntao, SHEN Ruonan, et al. Research progress on the effects of grazing on grassland ecosystem[J]. Chinese Journal of Plant Ecology, 2020, 44(5): 553−564. DOI: 10.17521/cjpe.2019.0314.
[4] 慕宗杰. 草地生态系统的保护及治理对策[J]. 畜牧与饲料科学, 2009, 30(2): 48−50.

MU Zongjie. Protection and control countermeasures of giassland ecosystem[J]. Animal Husbandry and Feed Science, 2009, 30(2): 48−50. DOI: 10.3969/j.issn.1672-5190.2009.02.022.
[5] CHEN Xiaolong, LI Xu, SHAN Xiaofeng, et al. The effects of sand-fixing agents and Trichoderma longibrachiatum on soil quality and alfalfa growth in wind-sand soil[J]. Agriculture, 2025, 15(23): 2463. DOI: 10.3390/agriculture15232463.
[6] 侯星辰, 鲁绍伟, 向昌林, 等. 不同母质温带草地植物群落多样性对人为干扰的响应[J]. 应用生态学报, 2022, 33(8): 2153−2160.

HOU Xingchen, LU Shaowei, XIANG Changlin, et al. Responses of plant community diversity to human disturbance in temperate grassland with different soil parent materials[J]. Chinese Journal of Applied Ecology, 2022, 33(8): 2153−2160. DOI: 10.13287/j.1001-9332.202208.015.
[7] 白永飞, 赵玉金, 王扬, 等. 中国北方草地生态系统服务评估和功能区划助力生态安全屏障建设[J]. 中国科学院院刊, 2020, 35(6): 675−689.

BAI Yongfei, ZHAO Yujin, WANG Yang, et al. Assessment of ecosystem services and ecological regionalization of grasslands support establishment of ecological security barriers in northern China[J]. Bulletin of the Chinese Academy of Sciences, 2020, 35(6): 675−689. DOI: 10.16418/j.issn.1000-3045.20200515003.
[8] 李禄军, 曾德慧, 于占源, 等. 氮素添加对科尔沁沙质草地物种多样性和生产力的影响[J]. 应用生态学报, 2009, 20(8): 1838−1844.

LI Lujun, ZENG Dehui, YU Zhanyuan, et al. Effects of nitrogen addition on grassland species diversity and productivity in Keerqin Sandy Land[J]. Chinese Journal of Applied Ecology, 2009, 20(8): 1838−1844. DOI: 10.13287/j.1001-9332.2009.0302.
[9] 郑伟, 董全民, 李世雄, 等. 放牧强度对环青海湖高寒草原群落物种多样性和生产力的影响[J]. 草地学报, 2012, 20(6): 1033−1038.

ZHENG Wei, DONG Quanmin, LI Shixiong, et al. Impact of grazing intensities on community biodiversity and productivity of alpine grassland in Qinghai Lake Region[J]. Acta Agrestia Sinica, 2012, 20(6): 1033−1038.
[10] GANGULI A C, O’ROURKE M E. How vulnerable are rangelands to grazing?[J]. Science, 2022, 378(6622): 834. DOI: 10.1126/science.add4278.
[11] XIANG Mingxue, WU Junxi, DUO L, et al. The effects of grazing and fencing on grassland productivity and diversity in alpine grassland ecosystem in the Tibetan highland[J]. Global Ecology and Conservation, 2023, 44: e02495. DOI: 10.1016/j.gecco.2023.e02495.
[12] BELLARD C, BERTELSMEIER C, LEADLEY P, et al. Impacts of climate change on the future of biodiversity[J]. Ecology Letters, 2012, 15(4): 365−377. DOI: 10.1111/j.1461-0248.2011.01736.x.
[13] BULLOCK J M, HILL B C, DALE M P, et al. An experimental study of the effects of sheep grazing on vegetation change in a species-poor grassland and the role of seedling recruitment into gaps[J]. Journal of Applied Ecology, 1994, 31(3): 493−507. DOI: 10.2307/2404445.
[14] 史志华, 刘前进, 张含玉, 等. 近十年土壤侵蚀与水土保持研究进展与展望[J]. 土壤学报, 2020, 57(5): 1117−1127.

SHI Zhihua, LIU Qianjin, ZHANG Hanyu, et al. Study on soil erosion and conservation in the past 10 years: progress and prospects[J]. Acta Pedologica Sinica, 2020, 57(5): 1117−1127. DOI: 10.11766/trxb202002240070.
[15] AKIYAMA T, KAWAMURA K. Grassland degradation in China: methods of monitoring, management and restoration[J]. Grassland Science, 2007, 53(1): 1−17. DOI: 10.1111/j.1744-697x.2007.00073.x.
[16] 晏和飘, 李文龙, 梁天刚, 等. 青藏高原退化高寒草地恢复对不同措施响应的Meta分析[J]. 草地学报, 2021, 29(S1): 190−198.

YAN Hepiao, LI Wenlong, LIANG Tiangang, et al. Effects of different restoration measures on degraded alpine grasslands of the Tibetan Plateau: a meta-analysis[J]. Acta Agrestia Sinica, 2021, 29(S1): 190−198. DOI: 10.11733/j.issn.1007-0435.2021.Z1.022.
[17] SUN J, CHENG G W, LI W P. Meta-analysis of relationships between environmental factors and aboveground biomass in the alpine grassland on the Tibetan Plateau[J]. Biogeosciences, 2013, 10(3): 1707−1715. DOI: 10.5194/bg-10-1707-2013.
[18] JING Zhaobin, CHENG Jimin, SU Jishuai, et al. Changes in plant community composition and soil properties under 3-decade grazing exclusion in semiarid grassland[J]. Ecological Engineering, 2014, 64: 171−178. DOI: 10.1016/j.ecoleng.2013.12.023.
[19] XIONG Dingpeng, SHI Peili, SUN Yinliang, et al. Effects of grazing exclusion on plant productivity and soil carbon, nitrogen storage in alpine meadows in northern Tibet, China[J]. Chinese Geographical Science, 2014, 24(4): 488−498. DOI: 10.1007/s11769-014-0697-y.
[20] SHANG Zhanhuan, CAO Jingjuan, GUO Ruiying, et al. Effect of enclosure on soil carbon, nitrogen and phosphorus of alpine desert rangeland[J]. Land Degradation & Development, 2017, 28(4): 1166−1177. DOI: 10.1002/ldr.2283.
[21] PATRICK L B, FRASER L H, KERSHNER M W. Large-scale manipulation of plant litter and fertilizer in a managed successional temperate grassland[J]. Plant Ecology, 2008, 197(2): 183−195. DOI: 10.1007/s11258-007-9369-7.
[22] 张艺斐, 王兴, 潘海珠, 等. 中国北方荒漠草原长期围栏封育恢复效应及其环境依赖性[J]. 生态学报, 2025, 45(17): 8529−8543.

ZHANG Yifei, WANG Xing, PAN Haizhu, et al. Long-term enclosure restoration effects and their environmental dependencies in desert steppes of northern China[J]. Acta Ecologica Sinica, 2025, 45(17): 8529−8543. DOI: 10.20103/j.stxb.202406061313.
[23] 江康威, 张青青, 王亚菲, 等. 放牧干扰下天山北坡中段植物功能群特征及其与土壤环境因子的关系[J]. 植物生态学报, 2024, 48(6): 701−718.

JIANG Kangwei, ZHANG Qingqing, WANG Yafei, et al. Characteristics of plant functional groups and the relationships with soil environmental factors in middle part of northern slope of Tianshan Mountains under different grazing intensities[J]. Chinese Journal of Plant Ecology, 2024, 48(6): 701−718. DOI: 10.17521/cjpe.2023.0225.
[24] 吴建波, 包晓影, 李洁, 等. 不同围封年限对典型草原群落及大针茅种群特征的影响[J]. 草地学报, 2010, 18(4): 490−495.

WU Jianbo, BAO Xiaoying, LI Jie, et al. Influence of fencing duration on community and population of Stipa grandis in a typical steppe[J]. Acta Agrectir Sinica, 2010, 18(4): 490−495. DOI: 10.3969/j.issn.1007-0435.2010.04.003.
[25] 彭珍, 肖红, 潘涛涛, 等. 模拟践踏和降雨对天祝高寒区垂穗披碱草有性繁殖的影响[J]. 中国草地学报, 2020, 42(2): 14−21.

PENG Zhen, XIAO Hong, PAN Taotao, et al. Effects of simulated trampling and rainfall on sexual reproduction of Elymus nutans in Tianzhu alpine meadow[J]. Chinese Journal of Grassland, 2020, 42(2): 14−21. DOI: 10.16742/j.zgcdxb.20190132.
[26] 施颖, 胡廷花, 高红娟, 等. 两种放牧模式下高寒草甸群落植被构成及稳定性特征[J]. 草业学报, 2019, 28(9): 1−10.

SHI Ying, HU Tinghua, GAO Hongjuan, et al. The community vegetation composition and stability characteristics of alpine meadow under two grazing modes[J]. Acta Prataculturae Sinica, 2019, 28(9): 1−10. DOI: 10.11686/cyxb2018571.
[27] 中华人民共和国农业农村部. NY/T 3647—2020 草食家畜羊单位换算[S]. 北京: 中国农业出版社, 2020.

Ministry of Agriculture and Rural Affairs of the People’s Republic of China. Conversion of Herbivore Livestock Unit: NY/T 3647−2020[S]. Beijing: China Agriculture Press, 2020.
[28] WHITTAKER R H. Evolution and measurement of species diversity[J]. Taxon, 1972, 21(2/3): 213−251.
[29] 马克平. 生物群落多样性的测度方法Ⅰα多样性的测度方法(上)[J]. 生物多样性, 1994, 2(3): 162−168.

MA Keping. The measurement method of biological community diversity Ⅰ The measurement method of α diversity (Ⅰ)[J]. Chinese Biodiversity, 1994, 2(3): 162−168. DOI: 10.17520/biods.1994027.
[30] 马克平, 刘玉明. 生物群落多样性的测度方法Ⅰα多样性的测度方法(下)[J]. 生物多样性, 1994, 2(4): 231−239.

MA Keping, LIU Yuming. The measurement method of biological community diversity Ⅰ The measurement method of α diversity (Ⅱ)[J]. Chinese Biodiversity, 1994, 2(4): 231−239. DOI: 10.17520/biods.1994038.
[31] 张钦弟, 卫伟, 陈利顶, 等. 黄土高原草地土壤水分和物种多样性沿降水梯度的分布格局[J]. 自然资源学报, 2018, 33(8): 1351−1362.

ZHANG Qindi, WEI Wei, CHEN Liding, et al. Spatial variation of soil moisture and species diversity patterns along a precipitation gradient in the grasslands of the Loess Plateau[J]. Journal of Natural Resources, 2018, 33(8): 1351−1362. DOI: 10.31497/zrzyxb.20170726.
[32] MAESTRE F T, QUERO J L, GOTELLI N J, et al. Plant species richness and ecosystem multifunctionality in global drylands[J]. Science, 2012, 335(6065): 214−218. DOI: 10.1126/science.1215442.
[33] HOU Dongjie, HE Weiming, LIU Changcheng, et al. Litter accumulation alters the abiotic environment and drives community successional changes in two fenced grasslands in Inner Mongolia[J]. Ecology and Evolution, 2019, 9(16): 9214−9224. DOI: 10.1002/ece3.5469.
[34] ZHANG Yiwen, CHANG Xiaofeng, ZHANG Yuwei, et al. Litter accumulation suppresses grass production but facilitates shrub expansion in a long-term fenced grassland[J]. Ecosphere, 2024, 15(4): e4844. DOI: 10.1002/ECS2.4844.
[35] 段敏杰, 高清竹, 万运帆, 等. 放牧对藏北紫花针茅高寒草原植物群落特征的影响[J]. 生态学报, 2010, 30(14): 3892−3900.

DUAN Minjie, GAO Qingzhu, WAN Yunfan, et al. Effect of grazing on community characteristics and species diversity of Stipa purpurea alpine grassland in Northern Tibet[J]. Acta Ecologica Sinica, 2010, 30(14): 3892−3900. DOI: 10.20103/j.stxb.2010.14.027.
[36] WANG Chao, TANG Yujia. A global meta-analyses of the response of multi-taxa diversity to grazing intensity in grasslands[J]. Environmental Research Letters, 2019, 14(11): 114003. DOI: 10.1088/1748-9326/ab4932.
[37] 王启基, 周兴民. 高寒矮嵩草草甸禾草种群的生长发育节律及环境适应性[J]. 植物生态学与地植物学学报, 1991, 15(2): 168−176.

WANG Qiji, ZHOU Xingmin. The growth rhythm of the grasses populations and their adaptability of environment in Kobresia humilis meadow[J]. Chinese Journal of Plant Ecology, 1991, 15(2): 168−176.
[38] 汪诗平, 王艳芬, 陈佐忠. 气候变化和放牧活动对糙隐子草种群的影响[J]. 植物生态学报, 2003, 27(3): 337−343.

WANG Shiping, WANG Yanfen, CHEN Zuozhong. Effect of climate change and grazing on populations of Cleistogenes squarrosa in Inner Mongolia steppe[J]. Acta Phytoecologica Sinica, 2003, 27(3): 337−343.
[39] FOSTER B L, GROSS K L. Partitioning the effects of plant biomass and litter on Andropogon gerardi in old-field vegetation[J]. Ecology, 1997, 78(7): 2091−2104. DOI: 10.2307/2265947.
[40] SUDING K N, LAVOREL S, CHAPIN III F S, et al. Scaling environmental change through the community-level: a trait-based response-and-effect framework for plants[J]. Global Change Biology, 2008, 14(5): 1125−1140. DOI: 10.1111/j.1365-2486.2008.01557.x.
[41] 徐来仙, 郭秋菊, 姚兰, 等. 凋落物物理阻隔对水杉幼苗出土和早期生长的影响[J]. 浙江农林大学学报, 2022, 39(5): 1018−1027.

XU Laixian, GUO Qiuju, YAO Lan, et al. Effect of litter physical barrier on emergence and early growth of Metasequoia glyptostroboides seedlings[J]. Journal of Zhejiang A&F University, 2022, 39(5): 1018−1027. DOI: 10.11833/j.issn.2095-0756.20210704.
[42] LETTS B, LAMB E G, MISCHKOLZ J M, et al. Litter accumulation drives grassland plant community composition and functional diversity via leaf traits[J]. Plant Ecology, 2015, 216(3): 357−370. DOI: 10.1007/s11258-014-0436-6.
[43] 宋明华, 刘丽萍, 陈锦, 等. 草地生态系统生物和功能多样性及其优化管理[J]. 生态环境学报, 2018, 27(6): 1179−1188.

SONG Minghua, LIU Liping, CHEN Jin, et al. Biology, multi-function and optimized management in grassland ecosystem[J]. Ecology and Environment Sciences, 2018, 27(6): 1179−1188. DOI: 10.16258/j.cnki.1674-5906.2018.06.025.
[44] 赵菲, 谢应忠, 马红彬, 等. 封育对典型草原植物群落物种多样性及土壤有机质的影响[J]. 草业科学, 2011, 28(6): 887−891.

ZHAO Fei, XIE Yingzhong, MA Hongbin, et al. Effects of enclosure on species diversity and soil organic matter of typical steppe[J]. Pratacultural Science, 2011, 28(6): 887−891. DOI: 10.3969/j.issn.1001-0629.2011.06.003.
[45] LOREAU M, NAEEM S, LNCHAUSTI P. Biodiversity and Ecosystem Functioning: Synthesis and Perspectives[M]. Oxford: Oxford University Press, 2002. DOI:10.1093/oso/9780198515708.001.0001.
[46] XIONG Shaojun, NILSSON C. The effects of plant litter on vegetation: a meta-analysis[J]. Journal of Ecology, 1999, 87(6): 984−994. DOI: 10.1046/j.1365-2745.1999.00414.x.
[47] BAGCHI S, RITCHIE M E. Herbivore effects on above- and belowground plant production and soil nitrogen availability in the Trans-Himalayan shrub-steppes[J]. Oecologia, 2010, 164(4): 1075−1082. DOI: 10.1007/s00442-010-1690-5.
[48] SIEPIELSKI A M, MCPEEK M A. On the evidence for species coexistence: a critique of the coexistence program[J]. Ecology, 2010, 91(11): 3153−3164. DOI: 10.1890/10-0154.1.
[49] 贺聪, 马永才, 字洪标. 不同围封年限对高寒草地植物群落的影响[J]. 草业学报, 2025, 34(11): 17−30.

HE Cong, MA Yongcai, ZI Hongbiao. Effects of the duration of grazing exclusion on plant communities in alpine grassland[J]. Acta Prataculturae Sinica, 2025, 34(11): 17−30. DOI: 10.11686/cyxb2024523.
[50] WU Xinwei, WANG Yichen, SUN Shucun. Long-term fencing decreases plant diversity and soil organic carbon concentration of the Zoige alpine meadows on the eastern Tibetan Plateau[J]. Plant and Soil, 2021, 458(1): 191−200. DOI: 10.1007/s11104-019-04373-7.
[51] 杨丽丽, 龚吉蕊, 王忆慧, 等. 内蒙古温带草原不同放牧强度和围栏封育对凋落物分解的影响[J]. 植物生态学报, 2016, 40(8): 748−759.

YANG Lili, GONG Jirui, WANG Yihui, et al. Effects of grazing intensity and grazing exclusion on litter decomposition in the temperate steppe of Nei Mongol, China[J]. Chinese Journal of Plant Ecology, 2016, 40(8): 748−759. DOI: 10.17521/cjpe.2016.0051.
[52] 陈亮, 王绪高. 生物多样性与森林生态系统健康的几个关键问题[J]. 生态学杂志, 2008, 27(5): 816−820.

CHEN Liang, WANG Xugao. Biodiversity and forest ecosystem health: some key questions[J]. Chinese Journal of Ecology, 2008, 27(5): 816−820. DOI: 10.13292/j.1000-4890.2008.0178.
[53] CHAI Qinglin, MA Zhanying, CHANG Xiaofeng, et al. Optimizing management to conserve plant diversity and soil carbon stock of semi-arid grasslands on the Loess Plateau[J]. CATENA, 2019, 172: 781−788. DOI: 10.1016/j.catena.2018.09.034.
[54] YUAN Z Y, JIAO F, LI Y H, et al. Anthropogenic disturbances are key to maintaining the biodiversity of grasslands[J]. Scientific Reports, 2016, 6: 22132. DOI: 10.1038/srep22132.