[1] 曾凡鹏, 迟光宇, 陈欣, 等. 辽东山区不同林龄落叶松人工林土壤-根系C∶N∶P生态化学计量特征[J]. 生态学杂志, 2016, 35(7): 1819 − 1825.

ZENG Fanpeng, CHI Guangyu, CHEN Xin, et al. The stoichiometric characteristics of C, N and P in soil and root of larch (Larix spp.) plantation at different stand ages in mountainous region of eastern Liaoning Province, China [J]. Chin J Ecol, 2016, 35(7): 1819 − 1825.
[2] 梅莉, 张卓文, 谷加存, 等. 水曲柳和落叶松人工林乔木层碳、氮储量及分配[J]. 应用生态学报, 2009, 20(8): 1791 − 1796.

MEI Li, ZHANG Zhuowen, GU Jiacun, et al. Carbon and nitrogen storages and allocation in tree layers of Fraxinus mandshurica and Larix gmelinii plantations [J]. Chin J Appl Ecol, 2009, 20(8): 1791 − 1796.
[3] 纪文婧, 程小琴, 韩海荣, 等. 不同林龄华北落叶松人工林生物量及营养元素分布特征[J]. 应用与环境生物学报, 2016, 22(2): 277 − 284.

JI Wenjing, CHENG Xiaoqin, HAN Hairong, et al. The biomass and nutrient distribution in Larix principis-ruppechtii Magyr plantations at different forest age [J]. Chin J Appl Environ Biol, 2016, 22(2): 277 − 284.
[4] 唐仕姗, 杨万勤, 殷睿, 等. 中国森林生态系统凋落叶分解速率的分布特征及其控制因子[J]. 植物生态学报, 2014, 38(6): 529 − 539.

TANG Shishan, YANG Wanqin, YIN Rui, et al. Spatial characteristics in decomposition rate of foliar litter and controlling factors in Chinese forest ecosystems [J]. Chin J Plant Ecol, 2014, 38(6): 529 − 539.
[5] 潘建平, 王华章, 杨秀琴. 落叶松人工林地力衰退研究现状与进展[J]. 东北林业大学学报, 1997, 25(2): 59 − 63.

PAN Jianping, WANG Huazhang, YANG Xiuqin. Research state and advance on soil degradation under Larch plantations [J]. J Northeast For Univ, 1997, 25(2): 59 − 63.
[6] 王理德, 王方琳, 郭春秀, 等. 土壤酶学硏究进展[J]. 土壤, 2016, 48(1): 12 − 21.

WANG Lide, WANG Fanglin, GUO Chunxiu, et al. Review: progress of soil enzymology [J]. Soils, 2016, 48(1): 12 − 21.
[7] 曹慧, 孙辉, 杨浩, 等. 土壤酶活性及其对土壤质量的指示研究进展[J]. 应用与环境生物学报, 2003, 9(1): 105 − 109.

CAO Hui, SUN Hui, YANG Hao, et al. A review soil enzyme activity and its indication for soil quality [J]. Chin J Appl Environ Biol, 2003, 9(1): 105 − 109.
[8]

PAZ-FERREIRO J, FU Shenglei, MWNDEZ A, et al. Interactive effects of biochar and the earthworm pontoscolex corethrurus on plant productivity and soil enzyme activities [J]. J Soil Sediment, 2014, 14(3): 483 − 494.
[9] 刘捷豹, 陈光水, 郭剑芬, 等. 森林土壤酶对环境变化的响应研究进展[J]. 生态学报, 2017, 37(1): 110 − 117.

LIU Jiebao, CHEN Guangshui, GUO Jianfen, et al. Advances in research on the responses of forest soil enzymes to environmental change [J]. Acta Ecol Sin, 2017, 37(1): 110 − 117.
[10]

HILL B H, ELONEN C M, SEIFERT L R, et al. Microbial enzyme stoichiometry and nutrient limitation in US streams and rivers [J]. Ecol Indic, 2012, 18: 540 − 551.
[11]

OLANDER L P, VITOUSEK P M. Regulation of soil phosphatase and chitinase activityby N and P availability [J]. Biogeochemistry, 2000, 49(2): 175 − 191.
[12]

SINSABAUGH R L, HILL B H, SHAH J J F. Ecoenzymatic stoichiometry of microbial organic nutrient acquisition in soil and sediment [J]. Nature, 2009, 462: 795 − 798.
[13] 吴秀芝, 阎欣, 王波, 等. 荒漠草地沙漠化对土壤-微生物-胞外酶化学计量特征的影响[J]. 植物生态学报, 2018, 42(10): 1022 − 1032.

WU Xiuzhi, YAN Xin, WANG Bo, et al. Effects of desertification on the C∶N∶P stoichiometry of soil, microbes, and extracellular enzymes in a desert grassland [J]. Chin J Plant Ecol, 2018, 42(10): 1022 − 1032.
[14] 袁萍, 周嘉聪, 张秋芳, 等. 中亚热带不同森林更新方式生态酶化学计量特征[J]. 生态学报, 2018, 38(18): 6741 − 6748.

YUAN Ping, ZHOU Jiacong, ZHANG Qiufang, et al. Patterns of ecoenzymatic stoichiometry in midsubtropical forest regeneration [J]. Acta Ecol Sin, 2018, 38(18): 6741 − 6748.
[15] 牛瑞龙, 高星, 徐福利, 等. 秦岭中幼林龄华北落叶松针叶与土壤的碳氮磷生态化学计量特征[J]. 生态学报, 2016, 36(22): 7384 − 7392.

NIU Ruilong, GAO Xing, XU Fuli, et al. Carbon,nitrogen,and phosphorus stoichiometric characteristics of soil and leaves from young and middle aged Larix principis-rupprechtii growing in a Qinling Mountain plantation [J]. Acta Ecol Sin, 2016, 36(22): 7384 − 7392.
[16]

ALLISON S D, VITOUSEK P M. Responses of extracellular enzymes to simple and complex nutrient inputs [J]. Soil Biol Biochem, 2004, 37(5): 937 − 944.
[17] 牛小云, 孙晓梅, 陈东升, 等. 辽东山区不同林龄日本落叶松人工林土壤微生物、养分及酶活性[J]. 应用生态学报, 2015, 26(9): 2663 − 2672.

NIU Xiaoyun, SUN Xiaomei, CHEN Dongsheng, et al. Soil microorganisms,nutrients and enzyme activity of Larix kaempferi plantation under different ages in mountainous region of eastern Liaoning Province,China [J]. Chin J Appl Ecol, 2015, 26(9): 2663 − 2672.
[18] 刘欣, 彭道黎, 邱新彩. 华北落叶松不同林型土壤理化性质差异[J]. 应用与环境生物学报, 2018, 24(4): 735 − 743.

LIU Xin, PENG Daoli, QIU Xincai. Differences in soil physicochemical properties between different forest types of Larix principis-rupprechtii [J]. Chin J Appl Environ Biol, 2018, 24(4): 735 − 743.
[19]

CHEN Guangcheng, GAO Min, PANG Bopeng, et al. Top-meter soil organic carbon stocks and sources in restored mangrove forests of different ages [J]. For Ecol Manage, 2018, 422: 87 − 94.
[20] 魏圣钊, 李林, 骆晓, 等. 不同连栽代次的巨桉(Eucalyptus grandis)人工林土壤酶活性及其与土壤理化性质的关系[J]. 应用与环境生物学报, 2019, 25(6): 1312 − 1318.

WEI Shengzhao, LI Lin, LUO Xiao, et al. Soil enzyme activities and their relationships to soil physicochemical properties in different successive rotation plantations of Eucalyptus grandis [J]. Chin J Appl Environ Biol, 2019, 25(6): 1312 − 1318.
[21] 邓娇娇, 周永斌, 殷有, 等. 油松和蒙古栎混交对土壤微生物群落功能多样性的影响[J]. 生态学杂志, 2017, 36(11): 3028 − 3035.

DENG Jiaojiao, ZHOU Yongbin, YIN You, et al. Effects of mixed Pinus tabuliformis and Quercus mongolica plantation on the functional diversity of soil microbial community [J]. Chin J Ecol, 2017, 36(11): 3028 − 3035.
[22] 刘旭军, 田慧霞, 程小琴, 等. 凋落物处理对不同林龄华北落叶松针阔混交林土壤磷组分的影响[J]. 生态学杂志, 2019, 38(10): 3024 − 3032.

LIU Xujun, TIAN Huixia, CHENG Xiaoqin, et al. Effects of litter manipulation on soil phosphorus fractions in Larix principis-rupprechtii conifer and broadleaved forests at different ages [J]. Chin J Ecol, 2019, 38(10): 3024 − 3032.
[23] 张雪. 根河林业局森林资源变化及其发展对策[D]. 呼和浩特: 内蒙古农业大学, 2018.

ZHANG Xue. Study on Forest Resources Change and Development Countermeasures of Genhe Forest Bureau[D]. Huhehaote: Inner Mongolia Agricultural University, 2018.
[24] 孙海滨, 王美莲, 张红星, 等. 大兴安岭森林火灾与气象因子相关性研究[J]. 内蒙古农业大学学报, 2012, 33(5/6): 87 − 90.

SUN Haibin, WANG Meilian, ZHANG Hongxing, et al. Correlation analysis betweeb forest fire and meteorological elements in daxinganling mountain [J]. J Inn Mong Agric Univ, 2012, 33(5/6): 87 − 90.
[25]

SAIYAA-CORK K R, SINSABAUGH R L, ZAK D R. The effects of long term nitrogen deposition on extracellular enzyme activity in an Acer saccharum forest soil [J]. Soil Biol Biochem, 2002, 34(9): 1309 − 1315.
[26] 鲁如坤. 土壤农业化学分析方法[M]. 北京: 中国农业科技出版社, 2000.
[27]

VANCE E D, BROOKES P C, JENKINSON D S. An extraction method for measuring soil microbial biomass C [J]. Soil Biol Biochem, 1987, 19(6): 703 − 707.
[28] 乔航, 莫小勤, 罗艳华, 等. 不同林龄油茶人工林土壤酶化学计量及其影响因素[J]. 生态学报, 2019, 39(6): 1887 − 1896.

QIAO Hang, MO Xiaoqin, LUO Yanhua, et al. Patterns of soil ecoenzymatic stoichiometry and its influencing factors during stand development in Camellia oleifera plantations [J]. Acta Ecol Sin, 2019, 39(6): 1887 − 1896.
[29] 史军, 刘纪远, 高志强, 等. 造林对土壤碳储量影响的研究[J]. 生态学杂志, 2005, 24(4): 410 − 416.

SHI Jun, LIU Jiyuan, GAO Zhiqiang, et al. A review on the influence of afforestation on soil carbon storage [J]. Chin J Ecol, 2005, 24(4): 410 − 416.
[30] 魏孝荣, 邵明安. 黄土高原小流域土壤pH、阳离子交换量和有机质分布特征[J]. 应用生态学报, 2009, 20(11): 2710 − 2715.

WEI Xiaorong, SHAO Ming’ an. Distribution characteristics of soil pH, CEC and organic matter in a small watershed of the Loess Plateau [J]. Chin J Appl Ecol, 2009, 20(11): 2710 − 2715.
[31]

MOLLA M A Z, CHOWDHURY A A, ISLAM A, et al. Microbial mineralization of organic phosphate in soil [J]. Plant Soil, 1984, 78(3): 393 − 399.
[32]

ALLISON V J, CONDRON L M, PELTZER D A, et al. Changes in enzyme activities and soil microbial community composition along carbon and nutrient gradients at the Franz Josef chronosequence, New Zealand [J]. Soil Biol Biochem, 2007, 39(7): 1770 − 1781.
[33] 陈立新, 段文标, 乔璐. 落叶松人工林根际与非根际土壤养分特征及酸度研究[J]. 水土保持学报, 2011, 25(3): 131 − 135.

CHEN Lixin, DUAN Wenbiao, QIAO Lu. Study on nutrition characteristics and acidity in rhizosphere and non-rhizosphere soils in Larch plantations [J]. J Soil Water Conserv, 2011, 25(3): 131 − 135.
[34] 张立欣, 段玉玺, 王博, 等. 库布齐沙漠不同人工固沙灌木林土壤微生物量与土壤养分特征[J]. 应用生态学报, 2017, 28(12): 3871 − 3880.

ZHANG Lixin, DUAN Yuxi, WANG Bo, et al. Characteristics of soil microorganisms and soil nutrients in different sand-fixation shrub plantations in Kubuqi Desert [J]. Chin J Appl Ecol, 2017, 28(12): 3871 − 3880.
[35] 赵娜, 孟平, 张劲松, 等. 华北低丘山地不同退耕年限刺槐人工林土壤质量评价[J]. 应用生态学报, 2014, 25(2): 351 − 358.

ZHAO Na, MENG Ping, ZHANG Jingsong, et al. Soil quality assessment of Robinia psedudoacia plantations with various ages in the Grain-for-Green Program in hilly area of North China [J]. Chin JAppl Ecol, 2014, 25(2): 351 − 358.
[36] 段益莉, 李继侠, 江强, 等. 长白山东坡不同海拔落叶松土壤微生物碳代谢及酶活性研究[J]. 生态环境学报, 2019, 28(4): 652 − 660.

DUAN Yili, LI Jixia, JIANG Qiang, et al. Soil microbial carbon metabolism and enzyme activity of Larix olgensis along an altitudinal gradient on the eastern slope of Changbai Mountain, Northeast China [J]. Ecol Environ Sci, 2019, 28(4): 652 − 660.
[37]

ZHANG Xinyu, DONG Wenyu, DAI Xiaoqin, et al. Responses of absolute and specific soil enzyme activities to long term additions of organic and mineral fertilizer [J]. Sci Total Environ, 2015, 536: 59 − 67.
[38]

RAIESI F, BEHESHTI A. Soil specific enzyme activity shows more clearly soil responses to paddy rice cultivation than absolute enzyme activity in primary forests of northwest Iran [J]. Appl Soil Ecol, 2014, 75: 63 − 70.
[39] 林诚, 王飞, 李清华, 等. 不同施肥制度对黄泥田土壤酶活性及养分的影响[J]. 中国土壤与肥料, 2009(6): 24 − 27.

LIN Cheng, WANG Fei, LI Qinghua, et al. Effects of different fertilizer application strategies on nutrients and enzymatic activities in yellow clayey soil [J]. Soil Fert Sci China, 2009(6): 24 − 27.
[40]

BLOOM A, CHAPIN I F S, MOONEY H. Resource limitation in plants--an economic analogy [J]. Ann Rev Ecol Syst, 1985, 16: 363 − 392.
[41]

TIAN Hanqin, CHEN Guangsheng, ZHANG Chi, et al. Pattern and variation of C∶N∶P ratios in China’ s soils: a synthesis of observational data [J]. Biogeochemistry, 2010, 98: 139 − 151.