[1] 贺金生, 韩兴国. 生态化学计量学: 探索从个体到生态系统的统一化理论[J]. 植物生态学报, 2010, 34(1): 2 − 6.

HE Jinsheng, HAN Xingguo. Ecological stoichiometry: searching for unifying principles from individuals to ecosystems [J]. Chinese Journal of Plant Ecology, 2010, 34(1): 2 − 6.
[2]

AGREN G I. The C∶N∶P stoichiometry of autotrophs: theory and observations [J]. Ecology Letters, 2004, 7(3): 185 − 191.
[3] 王霖娇, 汪攀, 盛茂银. 西南喀斯特典型石漠化生态系统土壤养分生态化学计量特征及其影响因素[J]. 生态学报, 2018, 38(18): 6580 − 6593.

WANG Linjiao, WANG Pan, SHENG Maoyin. Stoichiometric characteristics of soil nutrient elements and its influencing factors in typical karst rocky desertification ecosystems, Southwest China [J]. Acta Ecologica Sinica, 2018, 38(18): 6580 − 6593.
[4] 王亚娟, 陈云明, 孙亚荣, 等. 黄土丘陵区油松人工林植物器官-凋落物-土壤化学计量特征的季节变化[J]. 水土保持学报, 2022, 36(4): 350 − 356.

WANG Yajun, CHEN Yunming, SUN Yarong, et al. Seasonal variation of plant organ-litter-soil stoichiometry in Pinus tabulae formis plantation in Loess Hilly Region [J]. Journal of Soil and Water Conservation, 2022, 36(4): 350 − 356.
[5]

GÜSEWELL S. N∶P ratios in terrestrial plants: variation and functional significance [J]. New Phytologist, 2004, 164(2): 243 − 266.
[6]

HUA Fangyuan, BRUIJNZEEL L A, MELI P, et al. The biodiversity and ecosystem service contributions and trade-offs of forest restoration approaches[J/OL]. Science, 2022, 376(6595): 839[2022-05-15]. doi: 10.1126/science.abl4649.
[7]

CHEN Xinlin, CHEN H Y H, SEARLE E B, et al. Negative to positive shifts in diversity effects on soil nitrogen over time [J]. Nature Sustainability, 2020, 4(3): 225 − 232.
[8] 李非凡, 孙冰, 裴男才, 等. 粤北3种林分凋落叶-根系-土壤生态化学计量特征[J]. 浙江农林大学学报, 2020, 37(1): 18 − 26.

LI Feifan, SUN Bing, PEI Nancai, et al. Characteristics of litter-root-soil ecological stoichiometry of three forest stands in northern Guangdong [J]. Journal of Zhejiang A&F University, 2020, 37(1): 18 − 26.
[9] 尉剑飞, 王誉陶, 张翼, 等. 黄土高原典型草原植被及土壤化学计量对降水变化的响应[J]. 草地学报, 2022, 30(3): 532 − 543.

WEI Jianfei, WANG Yutao, ZHANG Yi, et al. Response of vegetation and soil stoichiometry to precipitation gradients of typical steppe in the Loess Plateau [J]. Acta Agrestia Sinica, 2022, 30(3): 532 − 543.
[10] 董廷发. 不同海拔云南松林土壤养分及其生态化学计量特征[J]. 生态学杂志, 2021, 40(3): 672 − 679.

DONG Tingfa. Soil nutrients and ecological stoichiometry in Pinus yunnanensis forests at different elevations [J]. Chinese Journal of Ecology, 2021, 40(3): 672 − 679.
[11] 余茂源. 云南松种质资源与遗传多样性研究进展[J]. 林业调查规划, 2011, 36(3): 39 − 42.

YU Maoyuan. Research advances of Pinus yunnanensis germplasm resources and genetic diversity [J]. Forest Inventory and Planning, 2011, 36(3): 39 − 42.
[12] 李艳琼, 黄玉清, 徐广平, 等. 桂林会仙喀斯特湿地芦苇群落土壤养分及微生物活性[J]. 生态学杂志, 2018, 37(1): 64 − 74.

LI Yanqiong, HUANG Yuqing, XU Guangping, et al. Characteristics of soil nutrients and microbial activities of reed vegetation in the Huixian karst wetland, Guilin, China [J]. Chinese Journal of Ecology, 2018, 37(1): 64 − 74.
[13] 张雨鉴, 王克勤, 宋娅丽, 等. 滇中亚高山森林植物叶-凋落叶-土壤生态化学计量特征[J]. 生态学报, 2020, 40(21): 7648 − 7658.

ZHANG Yujian, WANG Keqin, SONG Yali, et al. Ecological stoichiometry of leaf-litter-soil of alpine forest plants in central Yunnan, China [J]. Acta Ecologica Sinica, 2020, 40(21): 7648 − 7658.
[14] 陶慧敏, 孙宁骁, 温家豪, 等. 滇南喀斯特地区灌木群落和人工林土壤元素化学计量特征[J]. 生态学报, 2019, 39(24): 9119 − 9130.

TAO Huimin, SUN Ningxiao, WEN Jiahao, et al. Characteristics of soil stoichiometry in native shrub and plantation communities in karst regions of Yunnan Province [J]. Acta Ecologica Sinica, 2019, 39(24): 9119 − 9130.
[15]

DERENNE S, LARGEAU C. A review of some important families of refractory macromolecules: composition, origin, and fate in soils and sediments [J]. Soil Science, 2001, 166(11): 833 − 847.
[16] 张子琦, 焦菊英, 陈同德, 等. 拉萨河流域洪积扇不同植被类型的土壤化学计量特征[J]. 生态学报, 2022, 42(16): 6801 − 6815.

ZHANG Ziqi, JIAO Juying, CHEN Tongde, et al. Soil stoichiometry characteristics of different vegetation types in alluvial fans of the Lhasa River Basin[J]. Acta Ecologica Sinica, 2022, 42(16): 6801 − 6815.
[17] 吕金林, 闫美杰, 宋变兰, 等. 黄土丘陵区刺槐、辽东栎林地土壤碳、氮、磷生态化学计量特征[J]. 生态学报, 2017, 37(10): 3385 − 3393.

LÜ Jinlin, YAN Meijie, SONG Bianlan, et al. Ecological stoichiometry characteristics of soil carbon, nitrogen, and phosphorus in an oak forest and a black locust plantation in the loess hilly region [J]. Acta Ecologica Sinica, 2017, 37(10): 3385 − 3393.
[18] 许宇星, 王志超, 竹万宽, 等. 雷州半岛3种速生人工林下土壤生态化学计量特征[J]. 浙江农林大学学报, 2018, 35(1): 35 − 42.

XU Yuxing, WANG Zhichao, ZHU Wankuan, et al. Ecological stoichiometric characteristics of soil C, N, P, and K in three types of plantations on the Leizhou Peninsula [J]. Journal of Zhejiang A&F University, 2018, 35(1): 35 − 42.
[19]

LIAO Chang, LONG Chunyuan, ZHANG Qian, et al. Stronger effect of litter quality than micro-organisms on leaf and root litter C and N loss at different decomposition stages following a subtropical land use change [J]. Functional Ecology, 2021, 36(4): 896 − 907.
[20]

WEI Xiaorong, REICH P B, HOBBIE S E, et al. Legumes regulate grassland soil N cycling and its response to variation in species diversity and N supply but not CO2 [J]. Global Change Biology, 2019, 25(7): 2396 − 2409.
[21]

ZHOU Guoyi, XU Shang, CIAIS P, et al. Climate and litter C/N ratio constrain soil organic carbon accumulation [J]. National Science Review, 2019, 6(4): 746 − 757.
[22] 潘嘉琛, 刘超, 董智, 等. 黄泛沙地不同林龄杨树人工林土壤团聚体及有机碳特征[J]. 水土保持研究, 2022, 29(3): 25 − 30,37.

PAN Jiachen, LIU Chao, DONG Zhi, et al. Distribution characteristics of soil aggregates and soil organic carbon in Populus artificial forest with different forest ages in Yellow River Flood Plain [J]. Research of Soil and Water Conservation, 2022, 29(3): 25 − 30,37.
[23] 曾昭霞, 王克林, 刘孝利, 等. 桂西北喀斯特森林植物-凋落物-土壤生态化学计量特征[J]. 植物生态学报, 2015, 39(7): 682 − 693.

ZENG Zhaoxia, WANG Kelin, LIU Xiaoli, et al. Stoichiometric characteristics of plants, litter and soils in karst plant communities of northwest Guangxi [J]. Chinese Journal of Plant Ecology, 2015, 39(7): 682 − 693.
[24]

FENG Jiguang, HE Keyi, ZHANG Qiufang, et al. Changes in plant inputs alter soil carbon and microbial communities in forest ecosystems [J]. Global Change Biology, 2022, 28(10): 3426 − 3440.
[25] 吴鹏, 崔迎春, 赵文君, 等. 喀斯特森林植被自然恢复过程中土壤化学计量特征[J]. 北京林业大学学报, 2019, 41(3): 80 − 92.

WU Peng, CUI Yingchun, ZHAO Wenjun, et al. Characteristics of soil stoichiometric in natural restoration process of Maolan karst forest vegetation, southwestern China [J]. Journal of Beijing Forestry University, 2019, 41(3): 80 − 92.
[26] 杜家颖, 王霖娇, 盛茂银, 等. 喀斯特高原峡谷石漠化生态系统土壤C、N、P生态化学计量学特征[J]. 四川农业大学学报, 2017, 35(1): 45 − 51.

DU Jiaying, WANG Llinjiao, SHENG Maoyin, et al. Soil C, N and P stoichiometry of rocky desertification ecosystems in the karst plateau canyon area [J]. Journal of Sichuan Agricultural University, 2017, 35(1): 45 − 51.
[27]

PANG Danbo, WANG Genzhu, LI Guijing, et al. Ecological stoichiometric Ccharacteristics of two typical plantations in the karst ecosystem of southwestern China[J/OL]. Forests, 2018, 9(2)[2022-05-16]. doi:10.3390/f9020056.
[28] 刘娜, 喻理飞, 赵庆, 等. 喀斯特高原石漠化区次生林叶片-枯落物-土壤连续体碳氮磷生态化学计量特征[J]. 应用与环境生物学报, 2020, 26(3): 681 − 688.

LIU Na, YU Lifei, ZHAO Qing, et al. C∶N∶P stoichiometry of leaf-litter-soil continuum in secondary forests of the rocky desertification regions of the karst plateau [J]. Chinese Journal of Applied and Environmental Biology, 2020, 26(3): 681 − 688.
[29] 任悦, 高广磊, 丁国栋, 等. 沙地樟子松人工林叶片-枯落物-土壤氮磷化学计量特征[J]. 应用生态学报, 2019, 30(3): 743 − 750.

REN Yue, GAO Guanglei, DING Guodong, et al. Stoichiometric characteristics of nitrogen and phosphorus in leaf-litter-soil system of Pinus sylvestris var. mongolica plantations [J]. Chinese Journal of Applied Ecology, 2019, 30(3): 743 − 750.
[30] 吴丽芳, 王紫泉, 王妍, 等. 喀斯特高原不同石漠化程度土壤C、N、P化学计量特征和酶活性的关系[J]. 生态环境学报, 2019, 28(12): 2332 − 2340.

WU Lifang, WANG Ziquan, WANG Yan, et al. Relationship between soil C, N, P stoichiometric characteristics and enzyme activity in karst plateau soils with different degree of rocky desertification [J]. Ecology and Environment Sciences, 2019, 28(12): 2332 − 2340.
[31] 毕建华, 苏宝玲, 于大炮, 等. 辽东山区不同森林类型生态化学计量特征[J]. 生态学杂志, 2017, 36(11): 3109 − 3115.

BI Jianhua, SU Baoling, YU Dapao, et al. Ecological stoichiometry of different forest types in mountainous region of eastern Liaoning Province [J]. Chinese Journal of Ecology, 2017, 36(11): 3109 − 3115.
[32] 刘艳, 查同刚, 王伊琨, 等. 北京地区栓皮栎和油松人工林土壤团聚体稳定性及有机碳特征[J]. 应用生态学报, 2013, 24(3): 607 − 613.

LIU Yan, ZHA Tonggang, WANG Yikun, et al. Soil aggregate stability and soil organic carbon characteristics in Quercus variabilis and Pinus tabulaeformis plantations in Beijing area [J]. Chinese Journal of Applied Ecology, 2013, 24(3): 607 − 613.
[33] 余芹芹, 乔娜, 胡夏嵩, 等. 植物根-土复合体固坡力学效应及模型研究现状与进展[J]. 中国水土保持, 2011(7): 51 − 54,69.

YU Qinqin, QIAO Na, HU Xiasong, et al. Current status and progress of study on mechanical effect and model of plant root-soil complex for slope consolidation [J]. Soil and Water Conservation in China, 2011(7): 51 − 54,69.