[1] 赵其国, 黄国勤, 马艳芹. 中国南方红壤生态系统面临的问题及对策[J]. 生态学报, 2013, 33(24): 7615 − 7622.

ZHAO Qiguo, HUANG Guoqin, MA Yanqin. The problems in red soil ecosystem in southern of China and its countermeasures [J]. Acta Ecologica Sinica, 2013, 33(24): 7615 − 7622.
[2]

WU Qifeng, LIAN Ruiyuan, BAI Meixia, et al. Biochar co-application mitigated the stimulation of organic amendments on soil respiration by decreasing microbial activities in an infertile soil [J]. Biology and Fertility of Soils, 2021, 57(6): 793 − 807.
[3] 赵惠丽, 董金琎, 师江澜, 等. 秸秆还田模式对小麦-玉米轮作体系土壤有机碳固存的影响[J]. 土壤学报, 2021, 58(1): 213 − 224.

ZHAO Huili, DONG Jinjin, SHI Jianglan, et al. Effect of straw returning mode on soil organic carbon sequestration [J]. Acta Pedologica Sinica, 2021, 58(1): 213 − 224.
[4] 包建平, 袁根生, 董方圆, 等. 生物质炭与秸秆施用对红壤有机碳组分和微生物活性的影响[J]. 土壤学报, 2020, 57(3): 721 − 729.

BAO Jianping, YUAN Gensheng, DONG Fangyuan, et al. Effects of biochar application and straw returning on organic carbon fractionations and microbial activities in a red soil [J]. Acta Pedologica Sinica, 2020, 57(3): 721 − 729.
[5] 吴传敬, 郭剑芬, 许恩兰, 等. 采伐残余物不同处理方式对杉木幼林土壤有机碳组分和相关酶活性的影响[J]. 土壤学报, 2019, 56(6): 1504 − 1513.

WU Chuanjing, GUO Jianfen, XU Enlan, et al. Effects of logging residue on composition of soil carbon and activity of related enzymes in soil of a young Chinese fir plantation as affected by residue handling mode [J]. Acta Pedologica Sinica, 2019, 56(6): 1504 − 1513.
[6] 魏夏新, 熊俊芬, 李涛, 等. 有机物料还田对双季稻田土壤有机碳及其活性组分的影响[J]. 应用生态学报, 2020, 31(7): 2373 − 2380.

WEI Xiaxin, XIONG Junfen, LI Tao, et al. Effects of different organic amendments on soil organic carbon and its labile fractions in the paddy soil of a double rice cropping system [J]. Chinese Journal of Applied Ecology, 2020, 31(7): 2373 − 2380.
[7] 张聪, 慕平, 尚建明. 长期持续秸秆还田对土壤理化特性、酶活性和产量性状的影响[J]. 水土保持研究, 2018, 25(1): 92 − 98.

ZHANG Cong, MU Ping, SHANG Jianming. Effects of continuous returning corn straw on soil chemical properties, enzyme activities and yield trait [J]. Research of Soil and Water Conservation, 2018, 25(1): 92 − 98.
[8] 梅楠, 刘琳, 隋鹏祥, 等. 秸秆还田方式对土壤理化性质及玉米产量的影响[J]. 玉米科学, 2017, 25(6): 87 − 94.

MEI Nan, LIU Lin, SUI Pengxiang, et al. Effects of tillage and straw management on brown soil physical and chemical properties and maize yield [J]. Journal of Maize Sciences, 2017, 25(6): 87 − 94.
[9] 张英, 武淑霞, 雷秋良, 等. 不同类型粪肥还田对土壤酶活性及微生物群落的影响[J]. 土壤, 2022, 54(6): 1175 − 1184.

ZHANG Ying, WU Shuxia, LEI Qiuliang, et al. Effects of different manures on soil enzyme activity and microbial community [J]. Soils, 2022, 54(6): 1175 − 1184.
[10] 吴红, 刘海霞, 周明夏, 等. 羊粪还田对巨峰葡萄园土壤理化性质的影响[J]. 广东农业科学, 2018, 45(12): 32 − 37.

WU Hong, LIU Haixia, ZHOU Mingxia, et al. Effects of applying sheep manure on soil properties of kyoho vineyard [J]. Guangdong Agricultural Sciences, 2018, 45(12): 32 − 37.
[11] 韩召强, 陈效民, 曲成闯, 等. 生物质炭施用对潮土理化性状、酶活性及黄瓜产量的影响[J]. 水土保持学报, 2017, 31(6): 272 − 278.

HAN Zhaoqiang, CHEN Xiaomin, QU Chengchuang, et al. Effects of biochar application on soil physicochemical properties, enzyme activities and cucumber yield [J]. Journal of Soil and Water Conservation, 2017, 31(6): 272 − 278.
[12] 吴涛, 冯歌林, 曾珍, 等. 生物质炭对盆栽黑麦草生长的影响及机理[J]. 土壤学报, 2017, 54(2): 525 − 534.

WU Tao, FENG Gelin, ZENG Zhen, et al. Effect of biochar addition on ryegrass growth in a pot experiment and its mechanism [J]. Acta Pedologica Sinica, 2017, 54(2): 525 − 534.
[13]

LU Weiwei, DING Weixin, ZHANG Junhua, et al. Biochar suppressed the decomposition of organic carbon in a cultivated sandy loam soil: a negative priming effect [J]. Soil Biology and Biochemistry, 2014, 76: 12 − 21.
[14]

FANG Yunying, SINGH B P, SINGH B. Temperature sensitivity of biochar and native carbon mineralisation in biochar-amended soils [J]. Agriculture,Ecosystems &Environment, 2014, 191: 158 − 167.
[15]

LI Guanlin, KIM S J, HAN S H, et al. Precipitation affects soil microbial and extracellular enzymatic responses to warming [J]. Soil Biology and Biochemistry, 2018, 120: 212 − 221.
[16]

SINSABAUGH R L, HILL B H, SHAH J J F, et al. Ecoenzymatic stoichiometry of microbial organic nutrient acquisition in soil and sediment [J]. Nature, 2009, 462: 795 − 798.
[17]

ZHENG Mianhai, CHEN Hao, LI Dejun, et al. Substrate stoichiometry determines nitrogen fixation throughout succession in southern Chinese forests [J]. Ecology Letters, 2020, 23(2): 336 − 347.
[18] 左宜平, 张馨月, 曾辉, 等. 大兴安岭森林土壤胞外酶活力的时空动态及其对潜在碳矿化的影响[J]. 北京大学学报(自然科学版), 2018, 54(6): 1311 − 1324.

ZUO Yiping, ZHANG Xinyue, ZENG Hui, et al. Spatiotemporal dynamics of soil extracellular enzyme activity and its influence on potential mineralization rate of soil organic carbon in forests of Daxing’ an Mountain range [J]. Acta Scientiarum Naturalium Universitatis, 2018, 54(6): 1311 − 1324.
[19]

MOORHEAD D L, SINSABAUGH R L, HILL B H, et al. Vector analysis of ecoenzyme activities reveal constraints on coupled C, N and P dynamics [J]. Soil Biology &Biochemistry, 2016, 93: 1 − 7.
[20]

CUI Yongxing, FANG Linchuan, DENG Lei, et al. Patterns of soil microbial nutrient limitations and their roles in the variation of soil organic carbon across a precipitation gradient in an arid and semi-arid region [J]. Science of The Total Environment, 2019, 658: 1440 − 1451.
[21] 陈浩宁, 周怀平, 文永莉, 等. 长期不同施肥下褐土养分及酶活性的生态化学计量特征[J]. 植物营养与肥料学报, 2022, 28(6): 972 − 983.

CHEN Haoning, ZHOU Huaiping, WEN Yongli, et al. Ecological stoichiometric characteristics of soil nutrients and eco-enzymatic activities under different long-term fertilizations in a cinnamon soil [J]. Journal of Plant Nutrition and Fertilizers, 2022, 28(6): 972 − 983.
[22]

CHEN Zhe, JIN Penghui, WANG Hui, et al. Ecoenzymatic stoichiometry reveals stronger microbial carbon and nitrogen limitation in biochar amendment soils: a meta-analysis [J/OL]. The Science of the total environment, 2022, 838(3): 156532[2023-08-13]. doi: 10.1016/j.scitotenv.2022.156532.
[23] 鲁如坤. 土壤农业化学分析方法[M]. 北京: 中国农业科技出版社, 2000.

LU Rukun. The Analysis Method of Soil Agricultural Chemistry [M]. Beijing: China Agricultural Science and Technology Press, 2000.
[24]

ROVIRA P, VALLEJO V R. Labile and recalcitrant pools of carbon and nitrogen in organic matter decomposing at different depths in soil: an acid hydrolysis approach [J]. Geoderma, 2002, 107(1/2): 109 − 141.
[25] 陈小云, 郭菊花, 刘满强, 等. 施肥对红壤性水稻土有机碳活性和难降解性组分的影响[J]. 土壤学报, 2011, 48(1): 125 − 131.

CHEN Xiaoyun, GUO Juhua, LIU Manqiang, et al. Effects of fertilization on lability and recalcitrancy of organic carbon of red soil paddy soils [J]. Acta Pedologica Sinica, 2011, 48(1): 125 − 131.
[26]

SAIYA-CORK K R, SINSABAUGH R L, ZAK D R. The effects of long term nitrogen deposition on extracellular enzyme activity in an Acer saccharum forest soil [J]. Soil Biology & Biochemistry, 2002, 34(9): 1309 − 1315.
[27]

DEFOREST J L. The influence of time, storage temperature, and substrate age on potential soil enzyme activity in acidic forest soils using MUB-linked substrates and L-DOPA [J]. Soil Biology &Biochemistry, 2009, 41(6): 1180 − 1186.
[28]

DENG Lei, PENG Changhui, HUANG Chunbo, et al. Drivers of soil microbial metabolic limitation changes along a vegetation restoration gradient on the Loess Plateau, China [J]. Geoderma, 2019, 353: 188 − 200.
[29] 王改兰, 段建南, 贾宁凤, 等. 长期施肥对黄土丘陵区土壤理化性质的影响[J]. 水土保持学报, 2006, 20(4): 82 − 85, 89.

WANG Gailan, DUAN Jiannan, JIA Ningfeng, et al. Effects of long-term fertilizatton on soil physical and chemical property in Loess Hilly Area [J]. Journal of Soil and Water Conservation, 2006, 20(4): 82 − 85, 89.
[30] 徐阳春, 沈其荣, 冉炜. 长期免耕与施用有机肥对土壤微生物生物量碳、氮、磷的影响[J]. 土壤学报, 2002, 39(1): 83 − 90.

XU Yangchun, SHEN Qirong, RAN Wei. Effects of zero-tillage and application of manure on soil microbial biomass C, N and P after sixteen years of cropping [J]. Acta Pedologica Sinica, 2002, 39(1): 83 − 90.
[31]

WEI Zibiao, YING Hao, GUO Xiaowei, et al. Substitution of mineral fertilizer with organic fertilizer in maize systems: a meta-analysis of reduced nitrogen and carbon emissions [J/OL]. Agronomy, 2020, 10(8): 1149[2023-08-13]. doi: 10.3390/agronomy10081149.
[32]

BLAGODATSKAYA E, BLAGODATSKY S, ANDERSON T H, et al. Microbial growth and carbon use efficiency in the rhizosphere and root-rree soil [J/OL]. PLoS One, 2014, 9(4): e93282[2023-08-13]. doi: 10.1371/journal.pone.0093282.
[33]

LIU Yurong, DELGADO-BAQUERIZO M, WANG Juntao, et al. New insights into the role of microbial community composition in driving soil respiration rates [J]. Soil Biology &Biochemistry, 2018, 118: 35 − 41.
[34] 熊佰炼, 谭必勇. 生物质炭还田利用对土壤酶活性影响研究现状[J]. 遵义师范学院学报, 2017, 19(3): 106 − 110.

XIONG Bailian, TAN Biyong. On advances in biochar effects on soil enzyme activities [J]. Journal of Zunyi Normal College, 2017, 19(3): 106 − 110.
[35] 姚兰, 张焕朝, 胡立煌, 等. 黄山不同海拔植被带土壤活性有机碳、氮及其与酶活性的关系[J]. 浙江农林大学学报, 2019, 36(6): 1069 − 1076.

YAO Lan, ZHANG Huanchao, HU Lihuang, et al. Soil labile organic carbon and nitrogen and their relationship with enzyme activities in different vegetation zones along an altitudinal gradient on Mount Huangshan [J]. Journal of Zhejiang A&F University, 2019, 36(6): 1069 − 1076.
[36]

GUO Kangying, ZHAO Yingzhi, LIU Yang, et al. Pyrolysis temperature of biochar affects ecoenzymatic stoichiometry and microbial nutrient-use efficiency in a bamboo forest soil [J/OL]. Geoderma, 2020, 363: 114162[2023-08-13]. doi: 10.1016/j.geoderma.2019.114162.
[37]

BAILEY V L, FANSLER S J, SMITH J L, et al. Reconciling apparent variability in effects of biochar amendment on soil enzyme activities by assay optimization [J]. Soil Biology &Biochemistry, 2011, 43(2): 296 − 301.
[38] 田静, 盛茂银, 汪攀, 等. 西南喀斯特土地利用变化对植物凋落物-土壤C、N、P化学计量特征和土壤酶活性的影响[J]. 环境科学, 2019, 40(9): 4278 − 4286.

TIAN Jing, SHENG Maoyin, WANG Pan, et al. Influence of land use change on litter and soil C, N, P stoichiometric characteristics and soil enzyme activity in karst ecosystem, southwest China [J]. Environmental Science, 2019, 40(9): 4278 − 4286.
[39] 王博, 周志勇, 张欢, 等. 针阔混交林中兴安落叶松比例对土壤化学性质和酶化学计量比的影响[J]. 浙江农林大学学报, 2020, 37(4): 611 − 622.

WANG Bo, ZHOU Zhiyong, ZHANG Huan, et al. Effect of Larix gmelinii proportion on soil chemical properties and enzymatic stoichiometry in mixed coniferous and broad-leaved forest [J]. Journal of Zhejiang A&F University, 2020, 37(4): 611 − 622.
[40] 喻岚晖, 王杰, 廖李容, 等. 青藏高原退化草甸土壤微生物量、酶化学计量学特征及其影响因素[J]. 草地学报, 2020, 28(6): 1702 − 1710.

YU Lanhui, WANG Jie, LIAO Lirong, et al. Soil microbial biomass, enzyme activities and ecological stoichiometric characteristics and influencing factors along degraded meadows on the Qinghai-Tibet Plateau [J]. Acta Agrestia Sinica, 2020, 28(6): 1702 − 1710.
[41]

XU Zhiwei, YU Guirui, ZHANG Xinyu, et al. Soil enzyme activity and stoichiometry in forest ecosystems along the North-South Transect in eastern China (NSTEC) [J]. Soil Biology and Biochemistry, 2017, 104: 152 − 163.
[42] 赵娜, 王小利, 何进, 等. 有机肥替代化学氮肥对黄壤活性有机碳组分、酶活性及作物产量的影响[J/OL]. 环境科学. 2023-11-01[2023-12-01]. https://doi.org/10.13227/j.hjkx.202307222.

ZHAO Na, WANG Xiaoli, HE Jin, et al. Effects of replacing chemical nitrogen fertilizer with organic fertilizer on active organic carbon fractions, enzyme activities, and crop yield in yellow soil [J/OL]. Environmental Science, 2023-11-01 [2023-12-01]. https://doi.org/10.13227/j.hjkx.202307222.
[43]

SINSABAUGH R L. Phenol oxidase, peroxidase and organic matter dynamics of soil [J]. Soil Biology and Biochemistry, 2010, 42(3): 391 − 404.
[44]

CUI Yongxing, WANG Xia, ZHANG Xingchang, et al. Soil moisture mediates microbial carbon and phosphorus metabolism during vegetation succession in a semiarid region [J/OL]. Soil Biology and Biochemistry, 2020, 147: 107814[2023-07-13]. doi: 10.1016/j.soilbio.2020.107814.