[1] 赵波. 甘蓝型油菜矮秆基因定位、克隆及功能分析[D]. 武汉: 华中农业大学, 2017.

ZHAO Bo. Gentic Mapping, Cloning and Functional Analysis of Dwarf Genes in Brassica napus L.[D]. Wuhan: Huazhong Agricultural University, 2017.
[2] 张甜. 芥菜紫叶基因Bj.Pur定位及候选基因分析[D]. 武汉: 华中农业大学, 2017.

ZHANG Tian. Mapping and Candidate Gene Analysis of Bj.Pur, a Gene Controlling Purple Leaf in Brassica juncea[D]. Wuhan: Huazhong Agricultural University, 2017.
[3] 赖艳, 付秋实, 吕建春, 等. 一个新的薄皮甜瓜叶色突变体的生理特性及超微结构分析[J]. 四川农业大学学报, 2018, 36(3): 372 − 379.

LAI Yan, FU Qiushi, LÜ Jianchun, et al. Analysis of physiological characteristics and chloroplast ultrastructure of a new leaf color mutant in melon [J]. J Sichuan Agric Univ, 2018, 36(3): 372 − 379.
[4] 刘红艳, 周芳, 李俊, 等. 芝麻黄化突变体YL1的叶片解剖学及光合特性[J]. 作物学报, 2017, 43(12): 1856 − 1863.

LIU Hongyan, ZHOU Fang, LI Jun, et al. Anatomical structure and photosynthetic characteristics of a yellow leaf mutant YL1 in sesame(Sesamum indicum L.) [J]. Acta Agronomica Sin, 2017, 43(12): 1856 − 1863.
[5] 杨小苗, 吴新亮, 刘玉凤, 等. 一个番茄EMS叶色黄化突变体的叶绿素含量及光合作用[J]. 应用生态学报, 2018, 29(6): 1983 − 1989.

YANG Xiaomiao, WU Xinliang, LIU Yufeng, et al. Analysis of chlorophyll and photosynthesis of a tomato chlorophyll-deficient mutant induced by EMS [J]. Chin J Appl Ecol, 2018, 29(6): 1983 − 1989.
[6] 迟鸣雨. 青梗菜黄化突变体生理特性及转录组分析[D]. 沈阳: 沈阳农业大学, 2017.

CHI Mingyu. Transcriptome and Physiological Characterization Analysis of a Chlorosis Mutant of Pachoi[D]. Shenyang: Shenyang Agricultural University, 2017.
[7] 郭士伟, 张云华, 金永庆, 等. 小白菜(Brassica chinensis L.)黄苗突变体的叶绿素荧光特性栽[J]. 作物学报, 2003, 29(6): 958 − 960.

GUO Shiwei, ZHANG Yunhua, JIN Yongqing, et al. Characterization of chlorophyll fluorescence in a mutant of Brassica chinensis L. with xanthan seedling leaves [J]. Acta Agronomica Sin, 2003, 29(6): 958 − 960.
[8] 张琨, 刘志勇, 单晓菲, 等. 青梗菜黄化突变体pylm遗传特性分析[J]. 沈阳农业大学学报, 2017, 48(1): 1 − 8.

ZHANG Kun, LIU Zhiyong, SHAN Xiaofei, et al. Genetic analysis of a yellow mutant pylm in pakchoi [J]. J Shenyang Agric Univ, 2017, 48(1): 1 − 8.
[9] 杨冲, 张扬勇, 方智远, 等. 甘蓝叶色黄化突变体YL-1的光合生理特性及其叶绿体的超微结构[J]. 园艺学报, 2014, 41(6): 1133 − 1144.

YANG Chong, ZHANG Yangyong, FANG Zhiyuan, et al. Photosynthetic physiological characteristics and chloroplast ultrastructure of yellow leaf mutant YL-1 in cabbage [J]. Acta Hortic Sin, 2014, 41(6): 1133 − 1144.
[10] 杜江涛. 大白菜金黄叶色突变基因lcm2的克隆及鉴定[D]. 沈阳: 沈阳农业大学, 2018.

DU Jiangtao. Cloning and Identification of a Golden Leaf Gene lcm2 in Chinese Cabbage[D]. Shenyang: Shenyang Agricultural University, 2018.
[11] 侯爱琳. 大白菜叶片黄化突变基因lcm3的克隆与鉴定[D]. 沈阳: 沈阳农业大学, 2018.

HOU Ailin. Cloning and Identification of lcm3, a Leaf Chlorosis Mutantion Gene in Chinese Cabbage[D]. Shenyang: Shenyang Agricultural University, 2018.
[12]

ZHAO Hua, YU Lei, HUAI Zexun, et al. Mapping and candidate gene identification defining BnChd1-1, a locus involved in chlorophyll biosynthesis in Brassica napus [J]. Acta Physiol Plant, 2014, 36(4): 859 − 870.
[13] 董遵, 刘敬阳, 马红梅, 等. 甘蓝型油菜黄化(苗)突变体的叶绿素含量及超微结构[J]. 中国油料作物学报, 2000, 22(3): 27 − 29, 34.

DONG Zun, LIU Jingyang, MA Hongmei, et al. Chlorophyll contents and chloroplast ultrastructure of chlorophyll deficient mutant in B. napus [J]. Chin J Oil Crop Sci, 2000, 22(3): 27 − 29, 34.
[14] 陈艳丽. 甘蓝型油菜黄化突变体的基因定位[D]. 武汉: 华中农业大学, 2011.

CHEN Yanli. Genetic Mapping of the Yellow Mutant Gene in Brassca napus[D]. Wuhan: Huazhong Agricultural University, 2011.
[15]

ZHU Lixia, ZENG Xinhua, CHEN Yanli, et al. Genetic characterisation and fine mapping of a chlorophyll-deficient mutant (BnaC.ygl) in Brassica napus [J]. Mol Breed, 2014, 34(2): 603 − 614.
[16] 杨胜洪, 杜林方, 赵云, 等. 抽薹期叶绿素缺乏油菜突变体类囊体膜的研究[J]. 云南植物研究, 2001, 23(1): 97 − 104.

YANG Shenghong, DU Linfang, ZHAO Yun, et al. Study on the thylakoid membranes from a chlorophyll-deficient oilseed rape mutant at the bolting stage [J]. Acta Bot Yunnan, 2001, 23(1): 97 − 104.
[17]

FRICK G, SU Qingxiang, APEL K, et al. An Arabidopsis porB porC double mutant lacking light-dependent NADPH: protochlorophyllide oxidoreductases B and C is highly chlorophyll-deficient and developmentally arrested [J]. Plant J Cell Mol Biol, 2003, 35(2): 141 − 153.
[18]

BANG W Y, JEONG I S, KIM D W, et al. Role of Arabidopsis CHL27 protein for photosynthesis, chloroplast development and gene expression profiling [J]. Plant Cell Physiol, 2008, 49(9): 1350 − 1363.
[19]

PRIVAT I, HAKIMI M A, BUHOT L, et al. Characterization of Arabidopsis plastid sigma-like transcription factors SIG1, SIG2 and SIG3 [J]. Plant Mol Biol, 2003, 51(3): 385 − 399.
[20]

KUMAR A M, SÖLL D. Antisense HEMA1 RNA expression inhibits heme and chlorophyll biosynthesis in Arabidopsis [J]. Plant Physiol, 2000, 122(1): 49 − 56.
[21]

KOBAYASHI K, KONDO M, FUKUDA H, et al. Galactolipid synthesis in chloroplast inner envelope is essential for proper thylakoid biogenesis, photosynthesis, and embryogenesis [J]. Proc Nat Acad Sci, 2007, 104(43): 17216 − 17221.
[22] 白大勇. 拟南芥真叶白化突变体cfl1的基因克隆与初步功能分析[D]. 开封: 河南大学, 2013.

BAI Dayong. Map-based Cloning and Functional Analysis of Tue-Leaves-Etiolation Mutant cfl1 in Arabidopsis thaliana[D]. Kaifeng: Henan University, 2013.
[23] 肖华贵, 杨焕文, 饶勇, 等. 甘蓝型油菜黄化突变体的光合特性及叶绿素荧光参数分析[J]. 作物学报, 2013, 39(3): 520 − 529.

XIAO Huagui, YANG Huanwen, RAO Yong, et al. Photosynthetic characteristics and chlorophyll fluorescence kinetic parameters analyses of chlorophyll-reduced mutant in Brassica napus L. [J]. Acta Agronomica Sin, 2013, 39(3): 520 − 529.
[24]

CHANG C S J, WU M S H. COP1-Mediated degradation of BBX22/LZF1 optimizes seedling development in Arabidopsis [J]. Plant Physiol, 2011, 156(1): 228 − 239.
[25] 李玮, 于澄宇, 胡胜武. 芥菜型油菜叶片黄化突变体的初步研究[J]. 西北农林科技大学学报(自然科学版), 2007, 35(9): 79 − 82.

LI Wei, YU Chengyu, HU Shengwu. Primary investigation on a chlorsis mutant in Brassica juncea L. [J]. J Northwest A&F Univ Nat Sci Ed, 2007, 35(9): 79 − 82.
[26]

GAO Hongbo, SAGE T L, OSTERYOUNG K W. FZL, an FZO-like protein in plants, is a determinant of thylakoid and chloroplast morphology [J]. Proc Nat Acad Sci, 2006, 103(17): 6759 − 6764.
[27]

KIM Y K, LEE J Y, CHO H S, et al. Inactivation of organellar glutamyl- and seryl-trna synthetases leads to developmental arrest of chloroplasts and mitochondria in higher plants [J]. J Biol Chem, 2005, 280(44): 37098 − 37106.
[28] 赵云, 王茂林, 李江, 等. 幼叶黄化油菜(Brassica napus L.)突变体Cr3529叶绿体超微结构观察[J]. 四川大学学报(自然科学版), 2003, 40(5): 974 − 977.

ZHAO Yun, WANG Maolin, LI Jiang, et al. Observation of the chloroplast in chlorophyll-reduced seeding mutant Cr3529, Brassica napus L. [J]. J Sichuan Univ Nat Sci Ed, 2003, 40(5): 974 − 977.
[29] 吴砚农. 小白菜黄化突变体生理特性及遗传特性分析[D]. 杭州: 浙江农林大学, 2019.

WU Yannong. Analysis of Physiological Characteristics and Genetic Characteristics of Pakchoi Yellowing Mutant[D]. Hangzhou: Zhejiang A&F University, 2019.
[30]

AUSTIN J, WEBBER A N. Photosynthesis in Arabidopsis thaliana mutants with reduced chloroplast number [J]. Photosynth Res, 2005, 85(3): 373 − 384.
[31]

OKAZAKI K, KABEYA Y, SUZUKI K, et al. The PLASTID DIVISION1 and 2 components of the chloroplast division machinery determine the rate of chloroplast division in land plant cell differentiation [J]. Plant Cell, 2009, 21(6): 1769 − 1780.
[32] 牟钰. 白菜黄化突变基因py2的精细定位[D]. 沈阳: 沈阳农业大学, 2018.

MU Yu. Fine Mapping of py2, a Gene Referred to Yellow Leaf Mutant in Packoi (Brassica campestris L. ssp. chinensis)[D]. Shenyang: Shenyang Agricultural University, 2018.
[33] 方怡然, 薛立. 盐胁迫对植物叶绿素荧光影响的研究进展[J]. 生态科学, 2019, 38(3): 225 − 234.

FANG Yiran, XUE Li. Research advances in the effect of salt stress on plant chlorophyll fluorescence [J]. Ecol Sci, 2019, 38(3): 225 − 234.
[34]

TSANG E W T, YANG Jingyi, CHANG Qing, et al. Chlorophyll reduction in the seed of Brassica napus with a glutamate 1-semialdehyde aminotransferase antisense gene [J]. Plant Mol Biol, 2003, 51(2): 191 − 201.
[35] 田颖, 黄谦心, 刘海衡, 等. 芥菜型油菜黄化突变体L638-y的遗传及黄化基因gr1的分子标记[J]. 西北农林科技大学学报(自然科学版), 2012, 40(12): 90 − 96.

TIAN Ying, HUANG Qianxin, LIU Haiheng, et al. Imheritance of chlorophyll-deficient mutant L638-y in Brassica juncea L. and molecular markers for chlorophyll-deficient gene gr1 [J]. J Northwest A&F Univ Nat Sci Ed, 2012, 40(12): 90 − 96.
[36] 吴自明, 张欣, 万建民, 等. 叶绿素生物合成的分子调控[J]. 植物生理学报, 2008, 44(6): 1064 − 1070.

WU Ziming, ZHANG Xin, WAN Jianmin, et al. Molecular regulation of chlorophyll biosynthesis [J]. Plant Physiol Commun, 2008, 44(6): 1064 − 1070.
[37]

NAGATA N. Identification of a vinyl reductase gene for chlorophyll synthesis in Arabidopsis thaliana and implications for the evolution of prochlorococcus species [J]. Plant Cell, 2005, 17(1): 233 − 240.
[38] 孙捷音, 张年辉, 杜林方. 油菜叶绿素b减少突变体Cr3529叶绿素生物合成的研究[J]. 西北植物研究, 2007, 27(10): 1962 − 1966.

SUN Jieyin, ZHANG Nianhui, DU Linfang. Chlorophyll biosynthesis in a chlorophyll b-deficient oilseed rape mutant Cr3529 [J]. Acta Bot Boreal-Occident Sin, 2007, 27(10): 1962 − 1966.
[39] 吕明, 刘海衡, 毛虎德, 等. 芥菜型油菜黄化突变体叶片叶绿素合成代谢变化[J]. 西北植物学报, 2010, 30(11): 2177 − 2183.

LÜ Ming, LIU Haiheng, MAO Hude, et al. Changes of chlorophyll synthesis metabolism in chlorophyll-deficient mutant in Brassica juncea [J]. Acta Bot Boreal-Occident Sin, 2010, 30(11): 2177 − 2183.
[40]

PONTIER D, ALBRIEUX C, JOYARD J, et al. Knock-out of the magnesium protoporphyrin Ⅸ methyltransferase gene in Arabidopsis: effects on chloroplast development and on chloroplast-to-nucleus signaling [J]. J Biol Chem, 2007, 282(4): 2297 − 2304.
[41]

LARKIN R M. GUN4, a regulator of chlorophyll synthesis and intracellular signaling [J]. Science, 2003, 299(5608): 902 − 906.
[42]

KOBAYASHI K, FUJII S, SASAKI D, et al. Transcriptional regulation of thylakoid galactolipid biosynthesis coordinated with chlorophyll biosynthesis during the development of chloroplasts in Arabidopsis[J]. Front Plant Sci, 2014, 5(11): 272. doi: 10.3389/fpls.2014.00272.
[43]

HUANG Yishiuan, LI Hsoumin. Arabidopsis CHLI2 can substitute for CHLI1 [J]. Plant Physiol, 2009, 150(2): 636 − 645.
[44]

WATERS M T, LANGDALE J A. The making of a chloroplast [J]. EMBO J, 2009, 28(19): 2861 − 2873.
[45]

CHANG C S J, LI Y H, CHEN L T, et al. LZF1, a HY5-regulated transcriptional factor, functions in Arabidopsis de-etiolation [J]. Plant J, 2008, 54(2): 205 − 219.
[46] 张年辉, 杜林方, 赵云, 等. 叶绿素缺乏油菜突变体的LHCⅡ多肽组成、蛋白含量与cab基因转录研究[J]. 西北植物学报, 2004, 24(3): 484 − 487.

ZHANG Nianhui, DU Linfang, ZHAO Yun, et al. Study on the polypeptide composition and content of LHC Ⅱ and the cab gene transcription inchlorophyll-reduced mutant of oilseed rape seedlings [J]. Acta Bot Boreali-Occident Sin, 2004, 24(3): 484 − 487.
[47]

MIURA E, KATO Y, MATSUSHIMA R, et al. The balance between protein synthesis and degradation in chloroplasts determines leaf variegation in Arabidopsis yellow variegated mutants [J]. Plant Cell, 2007, 19(4): 1313 − 1328.
[48]

MOCHIZUKI N, BRUSSLAN J A, LARKIN R, et al. Arabidopsis genomes uncoupled 5(GUN5) mutant reveals the involvement of Mg-chelatase H subunit in plastid-to-nucleus signal transduction [J]. Proc Nat Acad Sci, 2001, 98(4): 2053 − 2058.
[49]

NAESTED H. Arabidopsis VARIEGATED 3 encodes a chloroplast-targeted, zinc-finger protein required for chloroplast and palisade cell development [J]. J Cell Sci, 2004, 117(20): 4807 − 4818.
[50]

SAKAMOTO W. Coordinated regulation and complex formation of Yellow Variegated1 and Yellow Variegated2, chloroplastic FtsH metalloproteases involved in the repair cycle of Photosystem Ⅱ in Arabidopsis thylakoid membranes [J]. Plant Cell, 2003, 15(12): 2843 − 2855.
[51]

SJOGREN L L E, STANNE T M, ZHENG B, et al. Structural and functional insights into the chloroplast ATP-dependent clp protease in Arabidopsis [J]. Plant Cell, 2006, 18(10): 2635 − 2649.
[52]

ZALTSMAN A, ORI N, ADAM Z. Two types of FtsH protease subunits are required for chloroplast biogenesis and Photosystem Ⅱ repair in Arabidopsis [J]. Plant Cell, 2005, 17(10): 2782 − 2790.