[1] OLSEN A N, ERNST H A, LEGGIO L L, et al. NAC transcription factors: structurally distinct, functionally diverse [J]. Trends Plant Sci, 2005, 10(2): 79 − 87.
[2] HIBARA K, TAKADA S, TASAKA M. CUC1 gene activates the expression of SAM-related genes to induce adventitious shoot formation [J]. Plant J, 2004, 36(5): 687 − 696.
[3] ZHANG Qian, LUO Fang, ZHONG Yu, et al. Modulation of NAC transcription factor NST1 activity by XYLEM NAC DOMAIN1 regulates secondary cell wall formation in Arabidopsis [J]. J Exp Bot, 2019, 71(4): 1449 − 1458.
[4] PITAKSARINGKARN W, MATSUOKA K, ASAHINA M, et al. XTH20 and XTH19 regulated by ANAC071 under auxin flow are involved in cell proliferation in incised Arabidopsis inflorescence stems [J]. Plant J, 2014, 80(4): 604 − 614.
[5] KANEDA T, TAGA Y, TAKAI R, et al. The transcription factor OsNAC4 is a key positive regulator of plant hypersensitive cell death [J]. EMBO J, 2009, 28(7): 926 − 936.
[6] JEONG J S, KIM Y S, BAEK K H, et al. Root-specific expression of OsNAC10 improves drought tolerance and grain yield in rice under field drought conditions [J]. Plant Physiol, 2010, 153(1): 185 − 197.
[7] CHEN Xu, LU Songchong, WANG Yaofeng, et al. OsNAC2 encoding a NAC transcription factor that affects plant height through mediating the gibberellic acid pathway in rice [J]. Plant J, 2015, 82(2): 302 − 314.
[8] LIU Yuanlong, KE Lili, WU Guizhi, et al. miR3954 is a trigger of phasiRNAs that affects flowering time in citrus [J]. Plant J, 2017, 92(2): 263 − 275.
[9] TRUPKINSA, ASTIGUETA F, BAIGORRIA A H, et al. Identification and expression analysis of NAC transcription factors potentially involved in leaf and petal senescence in Petunia hybrid[J]. Plant Science, 2019, 287: 110195. doi: 10.1016/j.plantsci.2019.110195.
[10] JIANG Guoxiang, LI Zhiwei, SONG Yunbo, et al. LcNAC13 physically interacts with LcR1MYB1 to coregulate anthocyanin biosynthesis-related genes during litchi fruit ripening[J]. Biomolecules, 2019, 9(4): 135. doi: 10.3390/biom9040135.
[11] van DOORN W G, van MEETEREN U. Flower opening and closure: a review [J]. J Exp Bot, 2003, 54(389): 1801 − 1812.
[12] van DOORN W G, KAMDEE C. Flower opening and closure: an update [J]. J Exp Bot, 2014, 65(20): 5749 − 5757.
[13] IRISH V F. The Arabidopsis petal: a model for plant organogenesis [J]. Trends Plant Sci, 2008, 13(8): 430 − 436.
[14] ZONIA L, MUNNIK T. Life under pressure: hydrostatic pressure in cell growth and function [J]. Trends Plant Sci, 2007, 12(3): 90 − 97.
[15] SABLOWSKI R, MEYEROWITZ E M. A homolog of NO APICAL MERISTEM is an immediate target of the floral homeotic genes APETALA3/PISTILLATA [J]. Cell, 1998, 92(1): 93 − 103.
[16] JIANG Xinqiang, ZHANG Changqing, LÜ Peitao, et al. RhNAC3, a stress-associated NAC transcription factor, has a role in dehydration tolerance through regulating osmotic stress-related genes in rose petals [J]. Plant Biotechnol J, 2014, 12(1): 38 − 48.
[17] 王英, 张超, 付建新, 等. 桂花花芽分化和花开放研究进展[J]. 浙江农林大学学报, 2016, 33(2): 340 − 347.

WANG Ying, ZHANG Chao, FU Jianxin, et al. Progress on flower bud differentiation and flower opening in Osmanthus fragrans [J]. J Zhejiang A&F Univ, 2016, 33(2): 340 − 347.
[18] ZHANG Chao, WANG Yiguang, FU Jianxin, et al. Transcriptomic analysis and carotenogenic gene expression related to petal coloration in Osmanthus fragrans‘Yanhong Gui’ [J]. Trees-Struct Funct, 2016, 30(4): 1207 − 1223.
[19] CHEN Chengjie, CHEN Hao, ZHANG Yi, et al. TBtools: an integrative toolkit developed for interactive analyses of big biological data [J]. Mol Plant, 2020, 13(8): 1194 − 1202.
[20] 付建新, 张超, 王艺光, 等. 桂花组织基因表达中荧光定量PCR内参基因的筛选[J]. 浙江农林大学学报, 2016, 33(5): 727 − 733.

FU Jianxin, ZHANG Chao, WANG Yiguang, et al. Reference gene selection for quantitativereal-time polymerase chain reaction(qRT-PCR) normalization in the gene expression of sweet osmanthus tissues [J]. J Zhejiang A&F Univ, 2016, 33(5): 727 − 733.
[21] SOUER E, van HOUWELINGEN A, KLOOS D, et al. The no apical meristem gene of Petunia is required for pattern formation in embryos and flowers and is expressed at meristem and primordia boundaries [J]. Cell, 1997, 85(2): 159 − 170.
[22] NURUZZAMAN M, MANIMEKALAI R, AKHTER S, et al. Genome-wide analysis of NAC transcription factor family in rice [J]. Gene, 2010, 465(1/2): 30 − 44.
[23] LIU Xingwang, WANG Ting, BARTHOLOMEW E, et al. Comprehensive analysis of NAC transcription factors and their expression during fruit spine development in cucumber (Cucumis sativus L.)[J]. Hortic Res, 2018, 5(1): 31. doi: 10.1038/s41438-018-0036-z.
[24] MIN Xueyang, JIN Xiaoyu, ZHANG Zhengshe, et al. Genome-wide identification of NAC transcription factor family and functional analysis of the abiotic stress-responsive genes in Medicago sativa L. [J]. J Plant Growth Regul, 2020, 39(1): 324 − 337.
[25] KIM S M, KIM S G, KIM Y S, et al. Exploring membrane-associated NAC transcription factors in Arabidopsis: implications for membrane biology in genome regulation [J]. Nucl Acids Res, 2007, 35(1): 203 − 213.
[26] FANG Yujie, YOU Jun, XIE Kabin, et al. Systematic sequence analysis and identification of tissue-specific or stress-responsive genes of NAC transcription factor family in rice [J]. Mol Genet Genomics, 2008, 280(6): 547 − 563.
[27] KIM S G, LEE A K, YOON H K, et al. A membrane-bound NAC transcription factor NTL8 regulates gibberellic acid-mediated salt signaling in Arabidopsis seed germination [J]. Plant J, 2008, 55(1): 77 − 88.
[28] 李建琴, 张娟, 王学臣, 等. 膜系留转录因子ANAC089在拟南芥开花诱导过程中起负调控作用[J]. 中国科学: 生命科学, 2010, 40(5): 408 − 417.

LI Jianqin, ZHANG Juan, WANG Xuechen, et al. A membrane-tethered transcription factor ANAC089 negatively regulates floral initiation in Arabidopsis thaliana [J]. Sci Sin Vitae, 2010, 40(5): 408 − 417.
[29] BALAZADEH S, SIDDIQUI H, ALLU A D, et al. A gene regulatory network controlled by the NAC transcription factor ANAC092/AtNAC2/ORE1 during salt-promoted senescence [J]. Plant J, 2010, 62(2): 250 − 264.
[30] PEI Haixia, MA Nan, TIAN Ji, et al. An NAC transcription factor controls ethylene-regulated cell expansion in flower petals [J]. Plant Physiol, 2013, 163(2): 775 − 791.
[31] 罗云, 张超, 付建新, 等. 桂花扩展蛋白基因家族的鉴定和表达分析[J]. 农业生物技术学报, 2017, 25(8): 1289 − 1299.

LUO Yun, ZHANG Chao, FU Jianxin, et al. Identification and expression analysis of expansin gene family in Osmanthus fragrans [J]. J Agric Biotechnol, 2017, 25(8): 1289 − 1299.
[32] DAI Fanwei, ZHANG Changqing, JIANG Xinqiang, et al. RhNAC2 and RhEXPA4 are involved in the regulation of dehydration tolerance during the expansion of rose petals [J]. Plant Physiol, 2012, 160(4): 2064 − 2082.
[33] SÁNCHEZ-MONTESINO R, BOUZA-MORCILLO L, MARQUEZ J, et al. A regulatory module controlling GA-mediated endosperm cell expansion is critical for seed germination in Arabidopsis [J]. Mol Plant, 2019, 12(1): 71 − 85.