[1] 牛保亮, 陈志成, 李铭杰, 等. 长期干旱降低暖温带天然林下草本层生物多样性[J]. 林业科学研究, 2025, 38(5): 73−81.

NIU Baoliang, CHEN Zhicheng, LI Mingjie, et al. Long-term drought reduced the biodiversity of understory herb layer in warm temperate natural forests[J]. Forest Research, 2025, 38(5): 73−81. DOI: 10.12403/j.1001-1498.20250020.
[2] CHEN Xinli, REICH P B, CHEN Xin, et al. Environmental change shapes understory plant diversity and dominance in boreal forests[J]. Nature Communications, 2025, 16: 10579. DOI: 10.1038/s41467-025-65633-y.
[3] WANG Ying, GUNINA A, REICH P B, et al. Nutrient cycles in transition: C∶N∶P stoichiometry by forest conversion to plantations[J]. Forest Ecology and Management, 2025, 593: 122874. DOI: 10.1016/j.foreco.2025.122874.
[4] 王晨阳, 毕华兴, 王武, 等. 黄土高原不同林分密度油松人工林林下灌木层和草本层组成及多样性差异特征[J]. 浙江农林大学学报, 2026-09-26. https://link.cnki.net/urlid/33.1370.S.20260929.0917.002.

WANG Chenyang, BI Huaxing, WANG Wu, et al. Differences in composition and diversity of understory shrub and herb layers in Pinus tabuliformis plantations with different stand densities on the Loess Plateau[J]. Journal of Zhejiang A&F University, 2026-09-26. https://link.cnki.net/urlid/33.1370.S.20260929.0917.002.
[5] 杜雨霜, 吴刘萍, 陈杰, 等. 崩岗生态修复不同人工林林下入侵植物和本土植物对群落稳定性的影响[J]. 生态学报, 2024, 44(4): 1588−1600.

DU Yushuang, WU Liuping, CHEN Jie, et al. Effects of understory invasive plants and native plants on community stability in Benggang ecological restoration of different artificial forests[J]. Acta Ecologica Sinica, 2024, 44(4): 1588−1600. DOI: 10.20103/j.stxb.202304010641.
[6] LE BAGOUSSE-PINGUET Y, SOLIVERES S, GROSS N, et al. Phylogenetic, functional, and taxonomic richness have both positive and negative effects on ecosystem multifunctionality[J]. Proceedings of the National Academy of Sciences of the United States of America, 2019, 116(17): 8419−8424. DOI: 10.1073/pnas.1815727116.
[7] 黄小荣. 广西马尾松林植物功能多样性与生产力的关系[J]. 生物多样性, 2018, 26(7): 690−700.

HUANG Xiaorong. Relationship between plant functional diversity and productivity of Pinus massoniana plantations in Guangxi[J]. Biodiversity Science, 2018, 26(7): 690−700. DOI: 10.17520/biods.2018092.
[8] BONGERS F J, SCHMID B, BRUELHEIDE H, et al. Functional diversity effects on productivity increase with age in a forest biodiversity experiment[J]. Nature Ecology & Evolution, 2021, 5(12): 1594−1603. DOI: 10.1038/s41559-021-01564-3.
[9] 王宇宁, 冯天骄, 孙龙, 等. 晋西黄土区典型人工林与天然林林下植被物种多样性差异及其影响因素[J]. 北京林业大学学报, 2025, 47(4): 103−116.

WANG Yuning, FENG Tianjiao, SUN Long, et al. Differences and influencing factors of understory vegetation species diversity between typical plantations and natural forests in the loess area of western Shanxi Province, Northern China[J]. Journal of Beijing Forestry University, 2025, 47(4): 103−116. DOI: 10.12171/j.1000−1522.20240228.
[10] OXBROUGH A, FILLOY J, ODDI J A, et al. Understorey forest management as a potential tool to promote taxonomic and functional diversity in Eucalyptus plantations[J]. Ecosystems, 2025, 28(4): 37. DOI: 10.1007/s10021-025-00978-5.
[11] 石慧, 王孝安, 郭华. 秦岭华北落叶松人工林群落结构及物种多样性[J]. 安徽农学通报, 2008, 14(15): 159−162, 204.

SHI Hui, WANG Xiaoan, GUO Hua. Community structure and species diversity of Larix principis-rupprechtii plantation in Qinling Mountains[J]. Anhui Agricultural Science Bulletin, 2008, 14(15): 159−162, 204. DOI: 10.16377/j.cnki.issn1007-7731.2008.15.086.
[12] 柴宗政, 王得祥, 郝亚中, 等. 秦岭中段华北落叶松人工林演替动态[J]. 林业科学, 2014, 50(2): 14−21.

CHAI Zongzheng, WANG Dexiang, HAO Yazhong, et al. Succession dynamics of Larix principis-rupprechtii plantation in intermediate section of Qinling Mountains[J]. Scientia Silvae Sinicae, 2014, 50(2): 14−21. DOI: 10.11707/j.1001-7488.20140203.
[13] 李敏敏, 魏天兴, 李信良, 等. 黄土区蔡家川流域刺槐人工林林下物种多样性[J]. 浙江农林大学学报, 2018, 35(2): 227−234.

LI Minmin, WEI Tianxing, LI Xinliang, et al. Species diversity in the understory of a Robinia pseudoacacia plantation in the Caijiachuan Watershed of the Loess Plateau[J]. Journal of Zhejiang A&F University, 2018, 35(2): 227−234. DOI: 10.11833/j.issn.2095-0756.2018.02.005.
[14] BUNESS V, SUNDQVIST M K, TUHIN ALI S, et al. Resource quantity and heterogeneity drive successional plant diversity in managed and unmanaged boreal forests[J]. Ecography, 2025, 2025(7): e07676. DOI: 10.1111/ecog.07676.
[15] 付立华, 张岩, 张菲, 等. 塞罕坝地区森林群落演替规律初探[J]. 安徽农学通报, 2022, 28(2): 65−67.

FU Lihua, ZHANG Yan, ZHANG Fei, et al. Preliminary study on the succession law of forest community in Saihanba Area[J]. Anhui Agricultural Science Bulletin, 2022, 28(2): 65−67. DOI: 10.16377/j.cnki.issn1007-7731.2022.02.046.
[16] XU Jie, XIAO Yu, XIE Gaodi, et al. Assessment of the benefit diffusion of windbreak and sand fixation service in National Key Ecological Function Areas in China[J]. Aeolian Research, 2021, 52: 100728. DOI: 10.1016/j.aeolia.2021.100728.
[17] 姚丹阳. 塞罕坝华北落叶松人工纯林和混交林林下植被多样性研究[J]. 防护林科技, 2018(12): 5−8.

YAO Danyang. Vegetation diversity of Larix principis-rupprechtii plantation in Saihanba Region[J]. Protection Forest Science and Technology, 2018(12): 5−8. DOI: 10.13601/j.issn.1005-5215.2018.12.002.
[18] 尹海龙, 张志伟, 宋伟, 等. 塞罕坝北曼甸林场森林生态功能等级评价[J]. 北京林业大学学报, 2023, 45(1): 89−98.

YIN Hailong, ZHANG Zhiwei, SONG Wei, et al. Grade evaluation of forest ecological function for Beimandian Forest Farm of Saihanba, Hebei Province of Northern China[J]. Journal of Beijing Forestry University, 2023, 45(1): 89−98. DOI: 10.12171/j.1000-1522.20210373.
[19] 葛兆轩, 苑美艳, 单博文, 等. 塞罕坝华北落叶松人工林不同经营模式效果评价[J]. 林业科学研究, 2020, 33(5): 38−47.

GE Zhaoxuan, YUAN Meiyan, SHAN Bowen, et al. Evaluation of management modes on Larix principis-rupprechtii plantations in Saihanba of Hebei Province, China[J]. Forest Research, 2020, 33(5): 38−47. DOI: 10.13275/j.cnki.lykxyj.2020.05.005.
[20] HUANG Kaichuan, ZHAO Wen, LI Junning, et al. Afforestation-induced shifts in soil bacterial diversity and community structure in the Saihanba Region[J]. Microorganisms, 2024, 12(3): 479. DOI: 10.3390/microorganisms12030479.
[21] 田地, 迟小龙, 石亮, 等. 塞罕坝地区优势造林树种叶片化学计量特征及其环境驱动[J]. 植物生态学报, 2026, 50(2): 362−373.

TIAN Di, CHI Xiaolong, SHI Liang, et al. Stoichiometric characteristics of dominant afforestation tree species and their environmental drivers in Saihanba Region[J]. Chinese Journal of Plant Ecology, 2026, 50(2): 362−373. DOI: 10.17521/cjpe.2025.0161.
[22] 方精云, 王襄平, 沈泽昊, 等. 植物群落清查的主要内容、方法和技术规范[J]. 生物多样性, 2009, 17(6): 533−548.

FANG Jingyun, WANG Xiangping, SHEN Zehao, et al. Methods and protocols for plant community inventory[J]. Biodiversity Science, 2009, 17(6): 533−548. DOI: 10.3724/SP.J.1003.2009.09253.
[23] HU Yukun, PAN Xu, LIU Xuyan, et al. Above- and belowground plant functional composition show similar changes during temperate forest swamp succession[J]. Frontiers in Plant Science, 2021, 12: 658883. DOI: 10.3389/fpls.2021.658883.
[24] DIAS R A, BASTAZINI V A G, GIANUCA A T, et al. The spatial scaling of multiple dimensions of functional diversity in habitat islands[J]. Functional Ecology, 2025, 39(11): 3164−3180. DOI: 10.1111/1365-2435.70158.
[25] LIU Libin, GUAN Xiaoyin, WANG Yunquan, et al. Intermediate anthropogenic disturbances boost taxonomic and phylogenetic diversities but reduce functional diversity in subtropical forests of Eastern China[J]. Plants, 2025, 14(16): 2529. DOI: 10.3390/plants14162529.
[26] 潘红林, 江怡航, 刘娜, 等. 不同林龄及区域杉木人工林林下植被多样性差异[J]. 林业科学研究, 2025, 38(6): 89−99.

PAN Honglin, JIANG Yihang, LIU Na, et al. Understory vegetation diversity in Chinese fir plantations across stand ages and regions[J]. Forest Research, 2025, 38(6): 89−99. DOI: 10.12403/j.1001-1498.20250024.
[27] ZHANG Dianye, PENG Yunfeng, LI Fei, et al. Trait identity and functional diversity co-drive response of ecosystem productivity to nitrogen enrichment[J]. Journal of Ecology, 2019, 107(5): 2402−2414. DOI: 10.1111/1365-2745.13184.
[28] LAI Jiangshan, ZOU Yi, ZHANG Shuang, et al. Glmm. hp: an R package for computing individual effect of predictors in generalized linear mixed models[J]. Journal of Plant Ecology, 2022, 15(6): 1302−1307. DOI: 10.1093/jpe/rtac096.
[29] 康义, 辛学兵, 裴顺祥, 等. 华北落叶松人工林林下物种多样性特征研究[J]. 西北林学院学报, 2025, 40(1): 23−31, 82.

KANG Yi, XIN Xuebing, PEI Shunxiang, et al. Undergrowth species diversity characteristics of Larix principis-rupprechtii plantation[J]. Journal of Northwest Forestry University, 2025, 40(1): 23−31, 82. DOI: 10.3969/j.issn.1001-7461.2025.01.03.
[30] 许驭丹, 董世魁, 李帅, 等. 植物群落构建的生态过滤机制研究进展[J]. 生态学报, 2019, 39(7): 2267−2281.

XU Yudan, DONG Shikui, LI Shuai, et al. Research progress on ecological filtering mechanisms for plant community assembly[J]. Acta Ecologica Sinica, 2019, 39(7): 2267−2281. DOI: 10.5846/stxb201804260946.
[31] SAUGIER B. Plant strategies, vegetation processes, and ecosystem properties[J]. Plant Science, 2001, 161(4): 813. DOI: 10.1016/s0168-9452(01)00469-1.
[32] CACCIANIGA M, LUZZARO A, PIERCE S, et al. The functional basis of a primary succession resolved by CSR classification[J]. Oikos, 2006, 112(1): 10−20. DOI: 10.1111/j.0030-1299.2006.14107.x.
[33] 戴楠, 王寅, 王梓静, 等. 植被恢复年限对塞罕坝华北落叶松人工林土壤碳氮磷储量的影响[J]. 北京林业大学学报, 2025, 47(5): 92−99.

DAI Nan, WANG Yin, WANG Zijing, et al. Effects of vegetation recovery years on soil carbon, nitrogen and phosphorus stocks of Larix gmelinii var. principis-rupprechtii plantations in Saihanba, Hebei Province of Northern China[J]. Journal of Beijing Forestry University, 2025, 47(5): 92−99. DOI: 10.12171/j.1000−1522.20240415.
[34] 于景金, 谷建才, 李校, 等. 塞罕坝落叶松人工林下植物多样性研究[J]. 林业资源管理, 2008(5): 90−94.

YU Jingjin, GU Jiancai, LI Xiao, et al. Study on plant diversity of the larch plantation in Saihanba[J]. Forest Resources Management, 2008(5): 90−94. DOI: 10.13466/j.cnki.lyzygl.2008.05.013.
[35] 黎露, 王永琪, 马姜明, 等. 南亚热带马尾松人工林近自然恢复过程中林下木本植物生态位动态[J]. 广西科学, 2021, 28(5): 499−510.

LI Lu, WANG Yongqi, MA Jiangming, et al. Niche dynamics of understory woody plants during the near- natural restoration of Pinus massoniana plantations in southern subtropics[J]. Guangxi Sciences, 2021, 28(5): 499−510. DOI: 10.13656/j.cnki.gxkx.20211203.001.
[36] 罗桂生, 张殿松, 黄拯, 等. 大岗山森林群落物种多样性和功能多样性的海拔格局特征[J/OL]. 中南林业科技大学学报, 2026-09-16. https://doi.org/10.14067/j.cnki.1673-923x.202510037.

LUO Guisheng, ZHANG Diansong, HUANG Zheng, et al. Altitudinal pattern characteristics of species diversity and functional diversity in forest communities of Dagangshan[J/OL]. Journal of Central South University of Forestry and Technology, 2026-09-16. https://doi.org/10.14067/j.cnki.1673-923x.202510037.
[37] ROSADO B H P, DE MATTOS E A. On the relative importance of CSR ecological strategies and integrative traits to explain species dominance at local scales[J]. Functional Ecology, 2017, 31(10): 1969−1974. DOI: 10.1111/1365-2435.12894.
[38] ZHOU Tiancai, HOU Ge, SUN Jian, et al. Degradation shifts plant communities from S- to R-strategy in an alpine meadow, Tibetan Plateau[J]. Science of the Total Environment, 2021, 800: 149572. DOI: 10.1016/j.scitotenv.2021.149572.
[39] 马文芳. 黄土丘陵区植被恢复对土壤碳、氮储量及团聚体分布的影响[D]. 杨凌: 西北农林科技大学, 2018.

MA Wenfang. Effects of Vegetation Restoration on Distribution of Soil Carbon, Nitrogen, and Dggregates in the Loess Hilly Region[D]. Yangling: Northwest A&F University, 2018. DOI:10.27409/d.cnki.gxbnu.2018.000373.
[40] 张泽凡, 张学珍. 黄土高原植被恢复对土壤物理参数的影响——基于已发表数据的荟萃分析[J]. 地理科学进展, 2021, 40(6): 1012−1025.

ZHANG Zefan, ZHANG Xuezhen. Effects of vegetation restoration on soil physical parameters on the Loess Plateau: a meta-analysis based on published data[J]. Progress in Geography, 2021, 40(6): 1012−1025. DOI: 10.18306/dlkxjz.2021.06.011.
[41] 拓行行, 李玉华, 俞瀚林, 等. 宁南山区华北落叶松林下草本层群落特征及其影响因素[J]. 生态学杂志, 2023, 42(10): 2449−2458.

TUO Hanghang, LI Yuhua, YU Hanlin, et al. Community characteristics and influencing factors of herbaceous layer under Larix gmelinii var. principis-rupprechtii forests in a south mountainous area of Ningxia[J]. Chinese Journal of Ecology, 2023, 42(10): 2449−2458. DOI: 10.13292/j.1000-4890.202310.013.
[42] 王梓静, 冯建, 李天雪, 等. 不同林龄华北落叶松人工林土壤水源涵养特征[J]. 河北师范大学学报(自然科学版), 2026, 50(2): 176−185.

WANG Zijing, FENG Jian, LI Tianxue, et al. Soil water conservation characteristics across a chronosequence of Larix gmelinii var. principis-rupprechtii plantations[J]. Journal of Hebei Normal University (Natural Science), 2026, 50(2): 176−185. DOI: 10.13763/j.cnki.jhebnu.nse.202604004.
[43] 刘浩栋, 王赫, 荣星星, 等. 尖峰岭陆均松群落功能性状与植物多样性关系的环境驱动机制[J]. 生态学报, 2026, 46(10): 5384−5395.

LIU Haodong, WANG He, RONG Xingxing, et al. Environmental filtering drives functional trait variation and mediates plant diversity patterns in Dacrydium pectinatum de Laubenfels communities of Jianfengling, Hainan Island, China[J]. Acta Ecologica Sinica, 2026, 46(10): 5384−5395. DOI: 10.20103/j.stxb.202503080506.