香榧僵果主要病原菌的分离鉴定及生物学特性

楼科勋; 楼君; 赖思思; 叶浩杰; 喻卫武; 方南寿; 胡渊渊; 楼科勋; 楼君; 赖思思; 叶浩杰; 喻卫武; 方南寿; 胡渊渊

doi:10.11833/j.issn.2095-0756.20250575

[1]

黎章矩, 戴文圣. 中国香榧[M]. 北京: 科学出版社, 2007.

LI Zhangju, DAI Wensheng. Torreya grandis of China[M]. Beijing: Science Press, 2007.

[2]

王哲, 吴诗涵, 索金伟, 等. 香榧僵果调查与解剖学观察[J]. 浙江农业科学, 2021, 62(10): 1950−1952, 2026.

WANG Zhe, WU Shihan, SUO Jinwei, et al. Investigation and anatomic observation of Torreya grandis aborted seeds[J]. Journal of Zhejiang Agricultural Sciences, 2021, 62(10): 1950−1952, 2026. DOI: 10.16178/j.issn.0528-9017.20211016.

[3]

成豪, 吴家胜, 马爽, 等. 树形调整对香榧成花和坐果的影响[J]. 林业科学, 2023, 59(11): 49−58.

CHENG Hao, WU Jiasheng, MA Shuang, et al. Effects of tree shape adjustment on flower formation and fruit setting of Torreya grandis cv. Merrillii[J]. Scientia Silvae Sinicae, 2023, 59(11): 49−58. DOI: 10.11707/j.1001-7488.LYKX20230104.

[4]

孙前军. 北京平谷区桃细菌性黑斑病与炭疽病病原鉴定及生防菌初筛[D]. 南阳: 南阳师范学院, 2024.

SUN Qianjun. Identification of Peach Bacterial Black Spot and Anthracnose Pathogens in Pinggu District, Beijing, and Initial Screening of Biocontrol Fungi [D]. Nanyang: Nanyang Normal Unisersity, 2024. DOI: 10.27850/d.cnki.gnysf.2024.000009.

[5]

王伟博, 李松刚, 胡美姣, 等. 无核荔枝僵果病的病原菌鉴定、入侵途径及快速检测技术[J]. 热带作物学报, 2024, 45(3): 602−608.

WANG Weibo, LI Songgang, HU Meijiao, et al. Identification, invasion pathway and rapid detection technology of new mummy fruit pathogen of seedless Litchi[J]. Chinese Journal of Tropical Crops, 2024, 45(3): 602−608. DOI: 10.3969/j.issn.1000-2561.2024.03.017.

[6]

王冬, 梁鹏, 刘晓菊. 苹果花腐病发病规律及防治措施[J]. 河北果树, 2009(1): 36.

WANG Dong, LIANG Peng, LIU Xiaoju. Occurrence regularity and control measures of apple flower rot[J]. Hebei Fruits, 2009(1): 36. DOI: 10.3969/j.issn.1006-9402.2009.01.022.

[7]

张强. 富士苹果霉心病病原菌侵染特性研究[D]. 杨凌: 西北农林科技大学, 2012.

ZHANG Qiang. Infection Characteristics of Pathogens Associated with Moldy Core of Fuji Apple[D]. Yangling: Northwest A&F University, 2012.

[8]

戴蓬博. 粉红聚端孢引致富士苹果心腐病的侵染特征及致病机制[D]. 杨凌: 西北农林科技大学, 2019.

DAI Pengbo. Infection Characteristics and Pathogenic Mechanisms of Apple Core Rot Caused by Trichothecium Roseum[D]. Yangling: Northwest A&F University, 2019.

[9]

叶浩杰. 香榧雌球花花腐病主要病原菌鉴定及其发生规律初探[D]. 杭州: 浙江农林大学, 2024.

Ye Haojie. Identification and Occurrence Regularity of the Main Pathogen for Female Cone Blight in Torreya grandis [D]. Hangzhou: Zhejiang A&F University, 2024.

[10]

ZHANG Yuan, NAN Mina, XUE Huali. Isolating, identifying, and analyzing the biological characteristics of pathogens causing postharvest disease in fresh Radix Astragali[J]. Horticulturae, 2023, 9(9): 1019. DOI: 10.3390/horticulturae9091019.

[11]

BENSCH K, GROENEWALD J Z, MEIJER M, et al. Cladosporium species in indoor environments[J]. Studies in Mycology, 2018, 89: 177−301. DOI:10.1016/j.simyco.2018.03.002.

[12]

PRASANNATH K, SHIVAS R G, GALEA V J, et al. Novel Botrytis and Cladosporium species associated with flower diseases of Macadamia in Australia[J]. Journal of Fungi, 2021, 7(11): 898. DOI: 10.3390/jof7110898.

[13]

PRASANNATH K, GALEA V J, AKINSANMI O A. Diversity and pathogenicity of species of Botrytis, Cladosporium, Neopestalotiopsis and Pestalotiopsis causing flower diseases of Macadamia in Australia[J]. Plant Pathology, 2023, 72(5): 881−899. DOI: 10.1111/ppa.13707.

[14]

陈思颖, 焦瑞莲, 李志强, 等. 新疆葡萄枝孢果腐病病原菌的鉴定及生物学特性[J]. 植物保护, 2022, 48(4): 180−187, 202.

CHEN Siying, JIAO Ruilian, LI Zhiqiang, et al. Identification and biological characteristics of Cladosporium pathogens causing grape fruits rot in Xinjiang[J]. Plant Protection, 2022, 48(4): 180−187, 202. DOI: 10.16688/j.zwbh.2021243.

[15]

龙安琪, 朱泽, 李强, 等. 浙贝母芽枝霉斑病病原菌鉴定、生物学特性及生物药剂防治效果研究[J]. 植物病理学报, 2023, 53(6): 1047−1055.

LONG Anqi, ZHU Ze, LI Qiang, et al. Identification biological characteristics, and inhibition efficacy of plant-derived antifungal agents of pathogen Cladosporium spp. causing mildew spot of Fritillaria thunbergii[J]. Acta Phytopathologica Sinica, 2023, 53(6): 1047−1055. DOI: 10.13926/j.cnki.apps.000838.

[16]

BENSCH K, BRAUN U, GROENEWALD J Z, et al. The genus Cladosporium[J]. Studies in Mycology, 2012, 72: 1−401. DOI: 10.3114/sim0003.

[17]

CARBONE I, KOHN L M. A method for designing primer sets for speciation studies in filamentous ascomycetes[J]. Mycologia, 1999, 91(3): 553−556. DOI: 10.1080/00275514.1999.12061051.

[18]

马文平, 倪志婧, 任贤, 等. ‘玉金香’甜瓜采后真菌病原物鉴定及侵染规律的研究[J]. 北方园艺, 2011(17): 36−40.

MA Wenping, NI Zhijing, REN Xian, et al. Study on latent infection pattern and identification of postharvest fungal diseases on ‘Yujinxiang’ melon fruit[J]. Northern Horticulture, 2011(17): 36−40.

[19]

黄欣阳. 中国辣椒叶斑病病原学及分子检测研究[D]. 沈阳: 沈阳农业大学, 2013.

HUANG Xinyang. The Etiology and Molecular Detection of a Leaf Spot on Pepper in China[D]. Shenyang: Shenyang Agricultural University, 2013.

[20]

BAISWAR P, CHANDRA S, BAG T K, et al. Cladosporium oxysporum on Prunus nepalensis in India[J]. Australasian Plant Disease Notes, 2011, 6(1): 3−6. DOI: 10.1007/s13314-011-0002-1.

[21]

WANG Siyao, WANG Yong, LI Yan. Cladosporium spp. (Cladosporiaceae) isolated from Eucommia ulmoides in China[J]. MycoKeys, 2022, 91: 151−168. DOI: 10.3897/mycokeys.91.87841.

[22]

GU Xin, LIU Wei, JI Kewen, et al. First reports of Cladosporium tenuissimum causing leaf spots on Hydrangea macrophylla in Anhui Province in China[J]. Plant Disease, 2024, 108(5): 1392. DOI: 10.1094/pdis-10-23-2261-pdn.

[23]

MIYAKE N, TSUKASA M, KOUNO T, et al. First report of sooty spot caused by Cladosporium tenuissimum on mature calyces of Cape gooseberry grown in greenhouse[J]. Journal of General Plant Pathology, 2022, 88(4): 282−284. DOI: 10.1007/s10327-022-01070-9.

[24]

ZHANG Yuanyuan, XU Hongyan, DING Jian, et al. First report of Cladosporium scabrellum causing stipe spot disease of Morchella importuna in China[J]. Plant Disease, 2024, 108(1): 225. DOI: 10.1094/pdis-09-22-2152-pdn.

[25]

CAMILETTI B X, LICHTEMBERG P S F, PAREDES J A, et al. Characterization, pathogenicity, and fungicide sensitivity of Alternaria isolates associated with preharvest fruit drop in California Citrus[J]. Fungal Biology, 2022, 126(4): 277−289. DOI: 10.1016/j.funbio.2022.02.003.

[26]

SAN MARTÍN J, LAGOS M J, RUIZ B, et al. Understanding the association of apical necrosis in flowers and fruits of walnut (Juglans regia L. ) with causal microorganisms under field conditions[J]. European Journal of Plant Pathology, 2023, 166(4): 445−462. DOI: 10.1007/s10658-023-02673-4.

[27]

ARMAND A, FALLAHI M, ABSALAN S, et al. Mango microcosm: unveiling the species diversity and pathogenic potential of fungi isolated from mango (Mangifera indica) in northern Thailand[J]. Fungal Biology, 2025, 129(3): 101552. DOI: 10.1016/j.funbio.2025.101552.

[28]

武崇高, 朱桐, 崔云龙, 等. 月季枯枝病病原物鉴定、生物学特性与防治药剂筛选[J/OL]. 浙江农林大学学报, 2025-07-14. https: //link.cnki.net/urlid/33.1370.s.20250711.1951.002.

WU Chonggao, ZHU Tong, CUI Yunlong, et al. Identification, characterization and fungicide screening of the causal pathogen of branch blight in Rosa chinensis[J/OL]. Journal of Zhejiang A&F University, 2025-07-14. https://link.cnki.net/urlid/33.1370.s.20250711.1951.002

[29]

项存悌, 常国彬, 赵海峰, 等. 极细枝孢生物学特性的研究[J]. 东北林业大学学报, 1989, 17(3): 38−41.

XIANG Cunti, CHANG Guobin, ZHAO Haifeng, et al. Study on biological characters of Cladosporium tenuissimum Cook[J]. Journal of Northeast Forestry University, 1989, 17(3): 38−41. DOI: 10.13759/j.cnki.dlxb.1989.03.006.

[30]

王树和, 周彩勤, 张奎望, 等. 金叶女贞棒孢叶斑病菌的生物学特性、致病性及系统发育[J]. 浙江农林大学学报, 2019, 36(6): 1174−1181.

WANG Shuhe, ZHOU Caiqin, ZHANG Kuiwang, et al. Characterization of Corynespora cassiicola from golden vicary privet based on phylogeny, biology, and pathogenicity[J]. Journal of Zhejiang A&F University, 2019, 36(6): 1174−1181. DOI: 10.11833/j.issn.2095-0756.2019.06.015.

[31]

周洁璐, 吴天昊, 巨云为, 等. 薄壳山核桃叶斑病病原菌生物学特性及室内药剂毒力测定[J]. 浙江农林大学学报, 2023, 40(5): 1018−1025.

ZHOU Jielu, WU Tianhao, JU Yunwei, et al. Biological characteristics and toxicity test of the pathogen of Carya illinoinensis leaf spot[J]. Journal of Zhejiang A&F University, 2023, 40(5): 1018−1025. DOI: 10.11833/j.issn.2095-0756.20230029.

[32]

ZHENG Fang, LI Lei, XIAO Xiaoe, et al. Species diversity of Cladosporium in Citrus and the genetic mechanisms for C. cladosporioides complex to adapt broad host plants[J]. Fungal Diversity, 2025, 133(1): 1−22. DOI: 10.1007/s13225-025-00559-w.

[33]

LIU Tao, ZHAO Xiaolong, ZHU Guangxia, et al. Shortage of storage carbohydrates mainly determines seed abscission in Torreya grandis ‘Merrillii’[J]. Horticultural Plant Journal, 2025, 11(2): 619−632. DOI: 10.1016/j.hpj.2023.10.007.

[34]

郝琪淳, 谢吉全, 戴文圣, 等. 香榧种实充实期叶面施肥对种实品质的影响[J]. 浙江农林大学学报, 2024, 41(3): 457−466.

HAO Qichun, XIE Jiquan, DAI Wensheng, et al. Effect of foliar fertilization on seed quality of Torreya grandis ‘Merrillii’ during seed filling period[J]. Journal of Zhejiang A& F University, 2024, 41(3): 457−466. DOI: 10.11833/j.issn.2095-0756.20230194.

[1]	黎章矩, 戴文圣. 中国香榧[M]. 北京: 科学出版社, 2007. LI Zhangju, DAI Wensheng. Torreya grandis of China[M]. Beijing: Science Press, 2007.
[2]	王哲, 吴诗涵, 索金伟, 等. 香榧僵果调查与解剖学观察[J]. 浙江农业科学, 2021, 62(10): 1950−1952, 2026. WANG Zhe, WU Shihan, SUO Jinwei, et al. Investigation and anatomic observation of Torreya grandis aborted seeds[J]. Journal of Zhejiang Agricultural Sciences, 2021, 62(10): 1950−1952, 2026. DOI: 10.16178/j.issn.0528-9017.20211016.
[3]	成豪, 吴家胜, 马爽, 等. 树形调整对香榧成花和坐果的影响[J]. 林业科学, 2023, 59(11): 49−58. CHENG Hao, WU Jiasheng, MA Shuang, et al. Effects of tree shape adjustment on flower formation and fruit setting of Torreya grandis cv. Merrillii[J]. Scientia Silvae Sinicae, 2023, 59(11): 49−58. DOI: 10.11707/j.1001-7488.LYKX20230104.
[4]	孙前军. 北京平谷区桃细菌性黑斑病与炭疽病病原鉴定及生防菌初筛[D]. 南阳: 南阳师范学院, 2024. SUN Qianjun. Identification of Peach Bacterial Black Spot and Anthracnose Pathogens in Pinggu District, Beijing, and Initial Screening of Biocontrol Fungi [D]. Nanyang: Nanyang Normal Unisersity, 2024. DOI: 10.27850/d.cnki.gnysf.2024.000009.
[5]	王伟博, 李松刚, 胡美姣, 等. 无核荔枝僵果病的病原菌鉴定、入侵途径及快速检测技术[J]. 热带作物学报, 2024, 45(3): 602−608. WANG Weibo, LI Songgang, HU Meijiao, et al. Identification, invasion pathway and rapid detection technology of new mummy fruit pathogen of seedless Litchi[J]. Chinese Journal of Tropical Crops, 2024, 45(3): 602−608. DOI: 10.3969/j.issn.1000-2561.2024.03.017.
[6]	王冬, 梁鹏, 刘晓菊. 苹果花腐病发病规律及防治措施[J]. 河北果树, 2009(1): 36. WANG Dong, LIANG Peng, LIU Xiaoju. Occurrence regularity and control measures of apple flower rot[J]. Hebei Fruits, 2009(1): 36. DOI: 10.3969/j.issn.1006-9402.2009.01.022.
[7]	张强. 富士苹果霉心病病原菌侵染特性研究[D]. 杨凌: 西北农林科技大学, 2012. ZHANG Qiang. Infection Characteristics of Pathogens Associated with Moldy Core of Fuji Apple[D]. Yangling: Northwest A&F University, 2012.
[8]	戴蓬博. 粉红聚端孢引致富士苹果心腐病的侵染特征及致病机制[D]. 杨凌: 西北农林科技大学, 2019. DAI Pengbo. Infection Characteristics and Pathogenic Mechanisms of Apple Core Rot Caused by Trichothecium Roseum[D]. Yangling: Northwest A&F University, 2019.
[9]	叶浩杰. 香榧雌球花花腐病主要病原菌鉴定及其发生规律初探[D]. 杭州: 浙江农林大学, 2024. Ye Haojie. Identification and Occurrence Regularity of the Main Pathogen for Female Cone Blight in Torreya grandis [D]. Hangzhou: Zhejiang A&F University, 2024.
[10]	ZHANG Yuan, NAN Mina, XUE Huali. Isolating, identifying, and analyzing the biological characteristics of pathogens causing postharvest disease in fresh Radix Astragali[J]. Horticulturae, 2023, 9(9): 1019. DOI: 10.3390/horticulturae9091019.
[11]	BENSCH K, GROENEWALD J Z, MEIJER M, et al. Cladosporium species in indoor environments[J]. Studies in Mycology, 2018, 89: 177−301. DOI:10.1016/j.simyco.2018.03.002.
[12]	PRASANNATH K, SHIVAS R G, GALEA V J, et al. Novel Botrytis and Cladosporium species associated with flower diseases of Macadamia in Australia[J]. Journal of Fungi, 2021, 7(11): 898. DOI: 10.3390/jof7110898.
[13]	PRASANNATH K, GALEA V J, AKINSANMI O A. Diversity and pathogenicity of species of Botrytis, Cladosporium, Neopestalotiopsis and Pestalotiopsis causing flower diseases of Macadamia in Australia[J]. Plant Pathology, 2023, 72(5): 881−899. DOI: 10.1111/ppa.13707.
[14]	陈思颖, 焦瑞莲, 李志强, 等. 新疆葡萄枝孢果腐病病原菌的鉴定及生物学特性[J]. 植物保护, 2022, 48(4): 180−187, 202. CHEN Siying, JIAO Ruilian, LI Zhiqiang, et al. Identification and biological characteristics of Cladosporium pathogens causing grape fruits rot in Xinjiang[J]. Plant Protection, 2022, 48(4): 180−187, 202. DOI: 10.16688/j.zwbh.2021243.
[15]	龙安琪, 朱泽, 李强, 等. 浙贝母芽枝霉斑病病原菌鉴定、生物学特性及生物药剂防治效果研究[J]. 植物病理学报, 2023, 53(6): 1047−1055. LONG Anqi, ZHU Ze, LI Qiang, et al. Identification biological characteristics, and inhibition efficacy of plant-derived antifungal agents of pathogen Cladosporium spp. causing mildew spot of Fritillaria thunbergii[J]. Acta Phytopathologica Sinica, 2023, 53(6): 1047−1055. DOI: 10.13926/j.cnki.apps.000838.
[16]	BENSCH K, BRAUN U, GROENEWALD J Z, et al. The genus Cladosporium[J]. Studies in Mycology, 2012, 72: 1−401. DOI: 10.3114/sim0003.
[17]	CARBONE I, KOHN L M. A method for designing primer sets for speciation studies in filamentous ascomycetes[J]. Mycologia, 1999, 91(3): 553−556. DOI: 10.1080/00275514.1999.12061051.
[18]	马文平, 倪志婧, 任贤, 等. ‘玉金香’甜瓜采后真菌病原物鉴定及侵染规律的研究[J]. 北方园艺, 2011(17): 36−40. MA Wenping, NI Zhijing, REN Xian, et al. Study on latent infection pattern and identification of postharvest fungal diseases on ‘Yujinxiang’ melon fruit[J]. Northern Horticulture, 2011(17): 36−40.
[19]	黄欣阳. 中国辣椒叶斑病病原学及分子检测研究[D]. 沈阳: 沈阳农业大学, 2013. HUANG Xinyang. The Etiology and Molecular Detection of a Leaf Spot on Pepper in China[D]. Shenyang: Shenyang Agricultural University, 2013.
[20]	BAISWAR P, CHANDRA S, BAG T K, et al. Cladosporium oxysporum on Prunus nepalensis in India[J]. Australasian Plant Disease Notes, 2011, 6(1): 3−6. DOI: 10.1007/s13314-011-0002-1.
[21]	WANG Siyao, WANG Yong, LI Yan. Cladosporium spp. (Cladosporiaceae) isolated from Eucommia ulmoides in China[J]. MycoKeys, 2022, 91: 151−168. DOI: 10.3897/mycokeys.91.87841.
[22]	GU Xin, LIU Wei, JI Kewen, et al. First reports of Cladosporium tenuissimum causing leaf spots on Hydrangea macrophylla in Anhui Province in China[J]. Plant Disease, 2024, 108(5): 1392. DOI: 10.1094/pdis-10-23-2261-pdn.
[23]	MIYAKE N, TSUKASA M, KOUNO T, et al. First report of sooty spot caused by Cladosporium tenuissimum on mature calyces of Cape gooseberry grown in greenhouse[J]. Journal of General Plant Pathology, 2022, 88(4): 282−284. DOI: 10.1007/s10327-022-01070-9.
[24]	ZHANG Yuanyuan, XU Hongyan, DING Jian, et al. First report of Cladosporium scabrellum causing stipe spot disease of Morchella importuna in China[J]. Plant Disease, 2024, 108(1): 225. DOI: 10.1094/pdis-09-22-2152-pdn.
[25]	CAMILETTI B X, LICHTEMBERG P S F, PAREDES J A, et al. Characterization, pathogenicity, and fungicide sensitivity of Alternaria isolates associated with preharvest fruit drop in California Citrus[J]. Fungal Biology, 2022, 126(4): 277−289. DOI: 10.1016/j.funbio.2022.02.003.
[26]	SAN MARTÍN J, LAGOS M J, RUIZ B, et al. Understanding the association of apical necrosis in flowers and fruits of walnut (Juglans regia L. ) with causal microorganisms under field conditions[J]. European Journal of Plant Pathology, 2023, 166(4): 445−462. DOI: 10.1007/s10658-023-02673-4.
[27]	ARMAND A, FALLAHI M, ABSALAN S, et al. Mango microcosm: unveiling the species diversity and pathogenic potential of fungi isolated from mango (Mangifera indica) in northern Thailand[J]. Fungal Biology, 2025, 129(3): 101552. DOI: 10.1016/j.funbio.2025.101552.
[28]	武崇高, 朱桐, 崔云龙, 等. 月季枯枝病病原物鉴定、生物学特性与防治药剂筛选[J/OL]. 浙江农林大学学报, 2025-07-14. https: //link.cnki.net/urlid/33.1370.s.20250711.1951.002. WU Chonggao, ZHU Tong, CUI Yunlong, et al. Identification, characterization and fungicide screening of the causal pathogen of branch blight in Rosa chinensis[J/OL]. Journal of Zhejiang A&F University, 2025-07-14. https://link.cnki.net/urlid/33.1370.s.20250711.1951.002
[29]	项存悌, 常国彬, 赵海峰, 等. 极细枝孢生物学特性的研究[J]. 东北林业大学学报, 1989, 17(3): 38−41. XIANG Cunti, CHANG Guobin, ZHAO Haifeng, et al. Study on biological characters of Cladosporium tenuissimum Cook[J]. Journal of Northeast Forestry University, 1989, 17(3): 38−41. DOI: 10.13759/j.cnki.dlxb.1989.03.006.
[30]	王树和, 周彩勤, 张奎望, 等. 金叶女贞棒孢叶斑病菌的生物学特性、致病性及系统发育[J]. 浙江农林大学学报, 2019, 36(6): 1174−1181. WANG Shuhe, ZHOU Caiqin, ZHANG Kuiwang, et al. Characterization of Corynespora cassiicola from golden vicary privet based on phylogeny, biology, and pathogenicity[J]. Journal of Zhejiang A&F University, 2019, 36(6): 1174−1181. DOI: 10.11833/j.issn.2095-0756.2019.06.015.
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[32]	ZHENG Fang, LI Lei, XIAO Xiaoe, et al. Species diversity of Cladosporium in Citrus and the genetic mechanisms for C. cladosporioides complex to adapt broad host plants[J]. Fungal Diversity, 2025, 133(1): 1−22. DOI: 10.1007/s13225-025-00559-w.
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