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小麦Triticum sestivum是中国最重要的粮食作物,小麦生产关系到国家粮食安全。麦田杂草对小麦生产造成了巨大的影响,如何有效地防控麦田杂草已成为农业生产中亟待解决的问题[1]。利用植物化感作用控制田间杂草被认为是环境友好型、可持续农业发展的新技术,已成为当代农业生态学研究中的重要领域之一[2-3]。诸多研究证明,小麦具有一定的化感潜力[4-9]。节节麦Aegilops tauschii为禾本科Poaceae山羊草属Aegilops 1年生或越年生植物,起源于东欧、西亚等地,是世界恶性杂草[10]。自1955年,节节麦首次在中国河南省新乡市被发现并采集到标本,如今已入侵中国陕西、山东、河北等地,并呈迅速蔓延的态势。由于节节麦与小麦亲缘关系较近,两者又具有类似的外部形态及生长习性,更重要的是节节麦拥有较强的分蘖、繁殖及适应性等特点,共同造成其成为麦田中最难防除的禾本科恶性杂草之一[11-12]。因严重威胁中国粮食生产安全,节节麦已被列入《中华人民共和国进境植物检疫性有害生物名录》。至今,未见有节节麦响应小麦化感作用的研究报道。为此,本研究以小麦不同部位水浸提液对节节麦种子萌发及幼苗生长的影响,揭示小麦化感作用机理,为麦田节节麦等杂草防控提供参考依据。
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随浸提液质量浓度的增加,节节麦种子发芽率呈先升后降的变化(图 1A)。浸提液质量浓度为5~10 g·L-1,不同部位水浸提液处理的种子发芽率均略有增加,但与对照差异均不显著(P>0.05);之后,发芽率均呈下降变化,至100 g·L-1时,与对照差异均达到显著水平(P<0.05),其中,茎叶水浸提液处理下降了50.0%,而根系处理则下降了45.0%。化感效应指数显示(图 1B):小麦不同部位水浸提液对节节麦种子萌发整体表现为“低促高抑”的化感效应。浸提液质量浓度为25~100 g·L-1时,对种子萌发的化感抑制作用为茎叶>根系。
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节节麦幼苗苗高随水浸提液质量浓度的增加呈先升后降的变化(图 2A),但小麦茎叶和根系不同质量浓度水浸提液的处理下,幼苗苗高与对照差异均不显著(P>0.05)。化感效应指数的变化显示(图 2B):小麦不同部位水浸提液对节节麦幼苗苗高呈“低促高抑”的化感效应。25~100 g·L-1茎叶水浸提液处理对幼苗苗高表现出质量浓度效应的化感抑制作用,且均明显大于根系水浸提液处理。
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节节麦幼苗根长随水浸提液质量浓度的增加整体呈先升后降的变化,茎叶和根系水浸提液质量浓度达50 g·L-1及以上时,幼苗根长较对照下降达到差异显著水平(图 3A,P<0.05)。化感效应指数显示(图 3B):小麦不同部位水浸提液对节节麦幼苗根长有“低促高抑”的化感作用。10~50 g·L-1质量浓度下对节节麦根长的化感抑制作用为茎叶<根系,而在100 g·L-1时,则为茎叶>根系。
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① 叶绿素质量分数。随水浸提液质量浓度的增加节节麦幼苗叶绿素质量分数整体呈下降趋势(图 4A)。低质量浓度(5~25 g·L-1)的水浸提液处理,叶绿素质量分数下降缓慢;水浸提液质量浓度达到50 g·L-1时,小麦不同部位水浸提液处理叶绿素质量分数与对照差异均达显著水平(P<0.05)。至100 g·L-1,小麦茎叶和根系水浸提液处理的叶绿素质量分数较对照分别下降了53.8%,48.4%。②相对电导率。随小麦水浸提液质量浓度的增加,节节麦幼苗相对电导率呈持续升高的趋势(图 4B),表明细胞膜透性不断增大。5~10 g·L-1范围内,不同部位水浸提液处理下相对电导率增加相对缓慢,与对照差异均未达到显著水平(P>0.05),之后,随水浸提液质量浓度的持续增加,相对电导率与对照差异均达显著水平(P<0.05)。至100 g·L-1,茎叶和根系水浸提液处理相对电导率分别较对照增加了2.1倍和1.8倍。③脯氨酸质量分数。随小麦水浸提液质量浓度的增加节节麦幼苗脯氨酸质量分数整体呈不断升高的趋势(图 4C)。茎叶和根系水浸提液处理中,脯氨酸质量分数与对照差异达到显著水平(P<0.05)的起始质量浓度分别为25 g·L-1,10 g·L-1。至100 g·L-1,茎叶和根系水浸提液处理脯氨酸质量分数分别较对照增加了68.2%,71.0%。④SOD活性。从图 4D可以看出:随小麦水浸提液质量浓度的增加节节麦幼苗SOD活性不断升高。茎叶和根系水浸提液处理中,SOD活性与对照差异达显著水平(P<0.05)的起始质量浓度均为10 g·L-1。50~100 g·L-1下,SOD活性增加缓慢;100 g·L-1与50 g·L-1浸提液处理下的SOD活性差异不显著(P>0.05)。⑤丙二醛质量摩尔浓度。随小麦水浸提液质量浓度的增加节节麦幼苗丙二醛质量摩尔浓度呈不断升高的趋势(图 4E)。5 g·L-1不同部位水浸提液处理下的丙二醛质量摩尔浓度与对照差异均不显著(P>0.05)。之后,随着水浸提液质量浓度的持续增加,节节麦幼苗丙二醛质量摩尔浓度不断升高,且与对照差异均达到显著水平(P<0.05)。至100 g·L-1,小麦茎叶和根系水浸提液处理丙二醛质量摩尔浓度较对照均增加了1.2倍。
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对节节麦幼苗苗高、根长以及叶绿素质量分数、相对电导率等生理指标进行综合分析,可反映小麦茎叶和根系水浸提液对节节麦幼苗的总体化感作用。从综合化感效应值可以看出(表 1):茎叶水浸提液对节节麦幼苗表现为化感抑制作用,而根系水浸提液则呈“低促高抑”的化感作用;化感抑制作用均随水浸提液质量浓度的增加而增大;相同水浸提液质量浓度条件下,茎叶水浸提液处理的化感效应值均大于或等于根系水浸提液,表明小麦水浸提液对节节麦幼苗的化感作用为茎叶≥根系。
表 1 小麦水浸提液对节节麦幼苗的综合化感效应
Table 1. Synthesis effects of aqueous extracts from wheat on t he seedlings of Aegilops tauschii
部位 ρ水浸提液/(g·L-1) IR 苗高 IR 根长 IRREC IR 叶绿素 IRSOD IRPro IRMDA 综合效应(Es) 茎叶 5 0.04 0.12 -0.28 -0.09 0.05 0.09 -0.07 -0.02 10 0.03 0.06 -0.25 -0.11 0.17 0.19 -0.27 -0.03 25 -0.11 -0.08 -0.48 -0.14 0.24 0.32 -0.41 -0.09 50 -0.16 -0.17 -0.62 -0.46 0.32 0.36 -0.48 -0.17 100 -0.19 -0.43 -0.68 -0.54 0.36 0.41 -0.55 -0.23 根系 5 0.06 0.01 -0.10 -0.03 0.09 0.11 -0.05 0.01 10 -0.08 -0.16 -0.12 -0.07 0.20 0.23 -0.21 -0.03 25 -0.09 -0.17 -0.43 -0.12 0.26 0.31 -0.33 -0.08 50 -0.11 -0.31 -0.55 -0.36 0.33 0.36 -0.42 -0.15 100 -0.16 -0.38 -0.65 -0.48 0.37 0.42 -0.54 -0.20
Seed germination and seedling growth of Aegilops tauschii with wheat extracts
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摘要: 为阐明小麦Triticum sestivum对入侵植物节节麦Aegilops tauschii的化感作用机制,采用培养皿生物测定法,依据萌发率、苗高、根长及超氧化物歧化酶(SOD)、丙二醛(MDA)等生理生化指标的变化,研究了小麦茎叶和根系水浸提液对节节麦种子萌发和幼苗生长的影响。结果表明:小麦茎叶和根系水浸提液对节节麦种子萌发、幼苗苗高和根长均表现出“低促高抑”的化感作用。生理生化指标结果显示,小麦不同部位水浸提液对节节麦幼苗生长表现出质量浓度效应的化感胁迫作用,随浸提液质量浓度的增加,节节麦幼苗相对电导率不断增加,叶绿素质量分数逐渐下降,当浸提液质量浓度达100 g·L-1时,与对照差异均达显著水平(P < 0.05)。此外,SOD活性、脯氨酸质量分数及丙二醛质量摩尔浓度均呈逐渐增加变化,其中,前期(0~5 g·L-1)丙二醛质量摩尔浓度较对照增加差异不显著(P>0.05),可能与节节麦自身调节有关,之后增加迅速,且较对照增加均达到差异显著水平(P < 0.05),表明节节麦幼苗生长受到小麦水浸提液化感胁迫作用的加剧。综合化感效应值显示,浸提液质量浓度为25~100 g·L-1时,小麦对节节麦幼苗化感抑制作用强弱为茎叶≥根系。Abstract: To clarify allelopathic potential and the preliminary mechanism of Triticum sestivum (wheat) on Aegilops tauschii, the effect of concentrations of T. sestivum extracts from stems, leaves, and roots on seed germination and seedling growth of A. tauschii were studied using Petri dish bioassays. Effects were based on physiological and ecological indexes, such as seed germination rate, seedling height, root length, superoxide dismutase (SOD) activity, malondialdehyde (MDA) content and other physiological and ecological index. Results showed that the aqueous extract increased membrane penetrability of A. tauschii, and enlarged the relative electrical conductivity, but decreased the chlorophyll mass fraction, reaching significance at 100 g·L-1 (P < 0.05). Also, water extracts increased SOD activities as well as proline and MDA content. At low concentrations (5 g·L-1), MDA increase was not significant (P>0.05), but with higher concentrations, MDA increased significantly (P < 0.05). According to the inhibition synthesis effect, the allelopathic inhibition of wheat stems and leaves was stronger than roots in the range of 25-100 g·L-1. Thus, seed germination along with seedling and root growth of A. tauschii were promoted at low concentrations of T. sestivum extracts and inhibited at high concentrations.
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Key words:
- plant physiology /
- allelopathy /
- Aegilops tauschii /
- seed germination /
- seedling growth
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表 1 小麦水浸提液对节节麦幼苗的综合化感效应
Table 1. Synthesis effects of aqueous extracts from wheat on t he seedlings of Aegilops tauschii
部位 ρ水浸提液/(g·L-1) IR 苗高 IR 根长 IRREC IR 叶绿素 IRSOD IRPro IRMDA 综合效应(Es) 茎叶 5 0.04 0.12 -0.28 -0.09 0.05 0.09 -0.07 -0.02 10 0.03 0.06 -0.25 -0.11 0.17 0.19 -0.27 -0.03 25 -0.11 -0.08 -0.48 -0.14 0.24 0.32 -0.41 -0.09 50 -0.16 -0.17 -0.62 -0.46 0.32 0.36 -0.48 -0.17 100 -0.19 -0.43 -0.68 -0.54 0.36 0.41 -0.55 -0.23 根系 5 0.06 0.01 -0.10 -0.03 0.09 0.11 -0.05 0.01 10 -0.08 -0.16 -0.12 -0.07 0.20 0.23 -0.21 -0.03 25 -0.09 -0.17 -0.43 -0.12 0.26 0.31 -0.33 -0.08 50 -0.11 -0.31 -0.55 -0.36 0.33 0.36 -0.42 -0.15 100 -0.16 -0.38 -0.65 -0.48 0.37 0.42 -0.54 -0.20 -
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https://zlxb.zafu.edu.cn/article/doi/10.11833/j.issn.2095-0756.2018.01.015